FNA: Erythranthe cordata vs. Erythranthe carsonensis

	Erythranthe cordata	Erythranthe carsonensis
	tinytooter monkeyflower	Carson Valley monkeyflower
Habit	Annuals, fibrous-rooted, sometimes producing leafy runners from basal nodes, stems often rooting at proximal nodes and appearing rhizomelike.	Annuals, taprooted, densely compact.
Stems	usually erect, usually simple, usually fistulose, 12–40(–100) cm, sparsely stipitate-glandular, hairs fine, gland-tipped.	erect, simple or branched from basal nodes, 2–7(–8) cm, minutely glandular, internodes shortened, not evident.
Leaves	basal and cauline, basal persistent; petiole: basal and proximals 6–20(–40) mm, midcauline to distals 0 mm; blade not connate, palmately 3–5(–7)-veined, orbicular to broadly elliptic-ovate or oblong-elliptic, cauline becoming broadly ovate to narrowly reniform, basal and mid cauline 15–30(–50) mm, gradually reduced in size distally to 6 mm, basal largest, distal closely paired, auriculate-subclasping, base cuneate to truncate or shallowly cordate, margins shallowly, evenly to unevenly dentate, apex obtuse to rounded, surfaces glabrous.	cauline, basal not persistent; petiole 0 mm; blade palmately 3-veined (in broader ones), linear to spatulate, (3–)5–23 × 1–5 mm, base truncate to truncate-cordate, clasping, margins entire, apex acute to obtuse, surfaces minutely glandular.
Flowers	plesiogamous, (5–)10–16, at distal nodes, in bracteate racemes, chasmogamous or cleistogamous.	herkogamous, 1–35, from distal or medial to distal nodes.
Styles	glabrous.	glabrous.
Corollas	yellow, red-spotted, abaxial limb deeper yellow, weakly bilaterally or radially symmetric, weakly bilabiate or regular; tube-throat sometimes tubular and not opening (cleistogamous), 8–14 mm, exserted 1–3 mm beyond calyx margin; limb not expanded or expanded 9–14 mm.	yellow, palate red-dotted and 1 large central spot, bilaterally symmetric, strongly bilabiate; tube-throat cylindric, distinct from abruptly expanding throat, (5–)7–11 mm, exserted beyond calyx margin; limb expanded 7–12(–15) mm, each lobe 2-fid, palate densely bearded.
Fruiting pedicels	10–30(–45) mm, longer than subtending leaves, minutely stipitate-glandular.	(3–)5–14 mm.
Fruiting calyces	nodding 45–90º, not red-dotted, broadly elliptic-ovoid, inflated, sagittally compressed, (8–)14–18(–20) mm, glabrous or sparsely stipitate-glandular to hirsutulous, sometimes mixed glandular-hirsutulous, throat closing, adaxial lobe not distinctly longer than abaxial, not falcate.	campanulate to widely urceolate, 4–7 mm, margins distinctly toothed or lobed, minutely glandular, ribs thickened, lobes pronounced, erect.
Capsules	included, stipitate, 5–7 mm.	included, 3–6 mm.
Anthers	included, glabrous.	included, glabrous.
2n	= 60.

	Erythranthe cordata	Erythranthe carsonensis
Phenology	Flowering (Jan–)Mar–Jun(–Nov).	Flowering Mar–Jun.
Habitat	Springs, seeps, stream edges, muddy banks, flood plains, marshes and swamps, wash bottoms, wet depressions, wet places among boulders.	Openings in sage brush/bitterbrush scrub in sand of decomposed granite.
Elevation	(600–)800–2400(–3000) m. ((2000–)2600–7900(–9800) ft.)	1400–1800 m. (4600–5900 ft.)
Distribution	AZ; CA; CO; NM; NV; TX; UT [WildflowerSearch map]	CA; NV
Discussion	Erythranthe cordata is characterized by its fibrous-rooted habit (annual in duration, without rhizomes but commonly rooting at the proximal nodes), short corollas and autogamous reproduction (anthers and stigma at the same level), closed calyces, sparsely villous-glandular vestiture (lacking hirtellous, eglandular hairs), and stems commonly fistulose in larger plants. The short corollas and other features of autogamous reproduction of E. cordata are diagnostic and prominent. Plants of E. cordata are highly variable in size, from tiny fibrous-rooted plants with nearly filiform stems to much larger individuals with fistulose stems rooting at proximal nodes. Erythranthe cordata and E. nasuta are sympatric in Arizona, southeastern New Mexico, and southern Utah, and small plants of each species may be similar in aspect, both with cleistogamous flowers and reduced vestiture. Erythranthe nasuta can be recognized by its distal and bracteal leaves with hirtellous to hirsutulous adaxial surfaces; a 10/x lens usually is required to see this feature, and it sometimes is most obvious around the leaf margins. The common name of Erythranthe cordata alludes to a fancied resemblance of the corollas to the horn of a diminutive trumpet. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Erythranthe carsonensis is restricted to the Carson Valley, Eagle Valley, and Washoe Valley region of Nevada and adjacent California, with one known disjunct occurrence about 58 km to the north in Nevada. The species was previously included in the broader concept of E. montioides but can be distinguished by its much branched and compact habit, linear to spatulate leaves with clasping bases, calyx with glabrous margins, and larger corolla with one large red spot in the center. Erythranthe carsonensis has been impacted by agriculture, urbanization, and other anthropogenic changes. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 422.	FNA vol. 17, p. 384.
Parent taxa	Phrymaceae > Erythranthe	Phrymaceae > Erythranthe
Sibling taxa	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms	Mimulus cordatus, M. maguirei	Mimulus rubellus var. latiflorus
Name authority	(Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012)	N. S. Fraga: Aliso 30: 59, figs. 17–21. (2012)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe carsonensis

tinytooter monkeyflower

Carson Valley monkeyflower

Habit

Annuals, fibrous-rooted, sometimes producing leafy runners from basal nodes, stems often rooting at proximal nodes and appearing rhizomelike.

Annuals, taprooted, densely compact.

Stems

usually erect, usually simple, usually fistulose, 12–40(–100) cm, sparsely stipitate-glandular, hairs fine, gland-tipped.

erect, simple or branched from basal nodes, 2–7(–8) cm, minutely glandular, internodes shortened, not evident.

Leaves

basal and cauline, basal persistent;

petiole: basal and proximals 6–20(–40) mm, midcauline to distals 0 mm;

blade not connate, palmately 3–5(–7)-veined, orbicular to broadly elliptic-ovate or oblong-elliptic, cauline becoming broadly ovate to narrowly reniform, basal and mid cauline 15–30(–50) mm, gradually reduced in size distally to 6 mm, basal largest, distal closely paired, auriculate-subclasping, base cuneate to truncate or shallowly cordate, margins shallowly, evenly to unevenly dentate, apex obtuse to rounded, surfaces glabrous.

cauline, basal not persistent;

petiole 0 mm;

blade palmately 3-veined (in broader ones), linear to spatulate, (3–)5–23 × 1–5 mm, base truncate to truncate-cordate, clasping, margins entire, apex acute to obtuse, surfaces minutely glandular.

Flowers

plesiogamous, (5–)10–16, at distal nodes, in bracteate racemes, chasmogamous or cleistogamous.

herkogamous, 1–35, from distal or medial to distal nodes.

Styles

glabrous.

Corollas

yellow, red-spotted, abaxial limb deeper yellow, weakly bilaterally or radially symmetric, weakly bilabiate or regular;

tube-throat sometimes tubular and not opening (cleistogamous), 8–14 mm, exserted 1–3 mm beyond calyx margin;

limb not expanded or expanded 9–14 mm.

yellow, palate red-dotted and 1 large central spot, bilaterally symmetric, strongly bilabiate;

tube-throat cylindric, distinct from abruptly expanding throat, (5–)7–11 mm, exserted beyond calyx margin;

limb expanded 7–12(–15) mm, each lobe 2-fid, palate densely bearded.

Fruiting pedicels

10–30(–45) mm, longer than subtending leaves, minutely stipitate-glandular.

(3–)5–14 mm.

Fruiting calyces

nodding 45–90º, not red-dotted, broadly elliptic-ovoid, inflated, sagittally compressed, (8–)14–18(–20) mm, glabrous or sparsely stipitate-glandular to hirsutulous, sometimes mixed glandular-hirsutulous, throat closing, adaxial lobe not distinctly longer than abaxial, not falcate.

campanulate to widely urceolate, 4–7 mm, margins distinctly toothed or lobed, minutely glandular, ribs thickened, lobes pronounced, erect.

Capsules

included, stipitate, 5–7 mm.

included, 3–6 mm.

Anthers

included, glabrous.

= 60.

Erythranthe cordata

Erythranthe carsonensis

Phenology

Flowering (Jan–)Mar–Jun(–Nov).

Flowering Mar–Jun.

Habitat

Springs, seeps, stream edges, muddy banks, flood plains, marshes and swamps, wash bottoms, wet depressions, wet places among boulders.

Openings in sage brush/bitterbrush scrub in sand of decomposed granite.

Elevation

(600–)800–2400(–3000) m. ((2000–)2600–7900(–9800) ft.)

1400–1800 m. (4600–5900 ft.)

Distribution

AZ; CA; CO; NM; NV; TX; UT

[WildflowerSearch map]

CA; NV

Discussion

Erythranthe cordata is characterized by its fibrous-rooted habit (annual in duration, without rhizomes but commonly rooting at the proximal nodes), short corollas and autogamous reproduction (anthers and stigma at the same level), closed calyces, sparsely villous-glandular vestiture (lacking hirtellous, eglandular hairs), and stems commonly fistulose in larger plants. The short corollas and other features of autogamous reproduction of E. cordata are diagnostic and prominent. Plants of E. cordata are highly variable in size, from tiny fibrous-rooted plants with nearly filiform stems to much larger individuals with fistulose stems rooting at proximal nodes.

Erythranthe cordata and E. nasuta are sympatric in Arizona, southeastern New Mexico, and southern Utah, and small plants of each species may be similar in aspect, both with cleistogamous flowers and reduced vestiture. Erythranthe nasuta can be recognized by its distal and bracteal leaves with hirtellous to hirsutulous adaxial surfaces; a 10/x lens usually is required to see this feature, and it sometimes is most obvious around the leaf margins.

The common name of Erythranthe cordata alludes to a fancied resemblance of the corollas to the horn of a diminutive trumpet.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe carsonensis is restricted to the Carson Valley, Eagle Valley, and Washoe Valley region of Nevada and adjacent California, with one known disjunct occurrence about 58 km to the north in Nevada. The species was previously included in the broader concept of E. montioides but can be distinguished by its much branched and compact habit, linear to spatulate leaves with clasping bases, calyx with glabrous margins, and larger corolla with one large red spot in the center. Erythranthe carsonensis has been impacted by agriculture, urbanization, and other anthropogenic changes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 422.

FNA vol. 17, p. 384.

Parent taxa

Phrymaceae > Erythranthe

Sibling taxa

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

Synonyms

Mimulus cordatus, M. maguirei

Mimulus rubellus var. latiflorus

Name authority

(Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012)

N. S. Fraga: Aliso 30: 59, figs. 17–21. (2012)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Erythranthe cordata

Erythranthe carsonensis

Erythranthe cordata

Erythranthe carsonensis