FNA: Erythranthe caespitosa vs. Erythranthe washingtonensis

	Erythranthe caespitosa	Erythranthe washingtonensis
	large mountain monkey-flower, mountain monkeyflower, Olympic monkeyflower, subalpine monkeyflower	John Day or Washington monkeyflower, Washington monkey-flower
Habit	Perennials, rhizomatous, rooting at proximal nodes, sometimes producing creeping, small-leaved runners, forming matted colonies, rhizomes filiform.	Annuals, fibrous-rooted or filiform-taprooted.
Stems	procumbent or decumbent to decumbent-ascending, delicate, usually in masses, terete or flattish, branched, 3–10 cm, glabrous, minutely hirtellous, or stipitate-glandular.	erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.
Leaves	basal and cauline; petiole: proximals 2–5 mm, distals 0 mm; blade often purple beneath, palmately 3-veined, orbicular to narrowly elliptic or ovate, proximals usually sublyrate, 3–12 mm, becoming larger distally, base cuneate to a short petiole, margins entire, mucronulate, or barely denticulate, apex obtuse, surfaces sparsely to moderately puberulent, hairs minute, stipitate-glandular.	cauline, basal not persistent; petiole 2–14 mm; blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.
Flowers	herkogamous, 1–3, from distal nodes, commonly solitary.	herkogamous, 1–6, from proximal to distal nodes.
Styles	minutely hirtellous.	hispid-hirtellous.
Corollas	yellow, dark red-spotted, bilaterally symmetric, bilabiate; tube-throat broadly funnelform to cylindric-funnelform, 15–18 mm, exserted beyond calyx margin; abaxial limb with deflexed-spreading lobes, adaxial with ascending lobes, palate partially closed.	yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 8–10 mm, exserted beyond calyx margin; limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate.
Fruiting pedicels	10–30(–40) mm, sparsely to moderately villous, hairs short, gland-tipped, sometimes hirtellous.	divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular.
Fruiting calyces	broadly campanulate, inflated, sagittally compressed, 7–15 mm, glabrous, minutely hirtellous, or stipitate-glandular, throat closing, proximalmost lobe pair upcurving, distalmost 3–5 mm, prominently protruding.	greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect.
Capsules	included, 4–5 mm.	included, 5–8.5 mm.
Anthers	included, glabrous.	included, glabrous.
2n		= 32.

	Erythranthe caespitosa	Erythranthe washingtonensis
Phenology	Flowering Jul–Sep.	Flowering May–Sep.
Habitat	Alpine meadows and slopes, stream banks, wet rocks in streams, wet crevices, talus.	Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps.
Elevation	1100–2000(–2300) m. (3600–6600(–7500) ft.)	700–1300 m. (2300–4300 ft.)
Distribution	WA; BC [WildflowerSearch map]	OR [WildflowerSearch map]
Discussion	Erythranthe caespitosa is endemic to northwestern and central Washington (Cascade and Olympic mountains) and adjacent British Columbia (Cascades, Selkirk Mountains and Chilliwack Valley, Coast Mountains). The plants have consistently small leaves with subentire margins, and the stems are consistently procumbent to decumbent-ascending, usually forming matted colonies. Erythranthe caespitosa and E. tilingii appear to be sympatric in counties of northwestern Washington, but this needs to be verified in the field. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 409.	FNA vol. 17, p. 396.
Parent taxa	Phrymaceae > Erythranthe	Phrymaceae > Erythranthe
Sibling taxa	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. willisii
Synonyms	Mimulus scouleri var. caespitosus, M. caespitosus, M. tilingii var. caespitosus	Mimulus washingtonensis
Name authority	(Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012)	(Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis)
Web links	Local floras: BC, OR, WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: OR, WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe washingtonensis

large mountain monkey-flower, mountain monkeyflower, Olympic monkeyflower, subalpine monkeyflower

John Day or Washington monkeyflower, Washington monkey-flower

Habit

Perennials, rhizomatous, rooting at proximal nodes, sometimes producing creeping, small-leaved runners, forming matted colonies, rhizomes filiform.

Annuals, fibrous-rooted or filiform-taprooted.

Stems

procumbent or decumbent to decumbent-ascending, delicate, usually in masses, terete or flattish, branched, 3–10 cm, glabrous, minutely hirtellous, or stipitate-glandular.

erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.

Leaves

basal and cauline;

petiole: proximals 2–5 mm, distals 0 mm;

blade often purple beneath, palmately 3-veined, orbicular to narrowly elliptic or ovate, proximals usually sublyrate, 3–12 mm, becoming larger distally, base cuneate to a short petiole, margins entire, mucronulate, or barely denticulate, apex obtuse, surfaces sparsely to moderately puberulent, hairs minute, stipitate-glandular.

cauline, basal not persistent;

petiole 2–14 mm;

blade palmately veined, deltate or ovate to ovate-lanceolate, 4–16(–23) × 2–11(–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped.

Flowers

herkogamous, 1–3, from distal nodes, commonly solitary.

herkogamous, 1–6, from proximal to distal nodes.

Styles

minutely hirtellous.

hispid-hirtellous.

Corollas

yellow, dark red-spotted, bilaterally symmetric, bilabiate;

tube-throat broadly funnelform to cylindric-funnelform, 15–18 mm, exserted beyond calyx margin;

abaxial limb with deflexed-spreading lobes, adaxial with ascending lobes, palate partially closed.

yellow with small reddish brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate;

tube-throat funnelform, 8–10 mm, exserted beyond calyx margin;

limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate.

Fruiting pedicels

10–30(–40) mm, sparsely to moderately villous, hairs short, gland-tipped, sometimes hirtellous.

divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular.

Fruiting calyces

broadly campanulate, inflated, sagittally compressed, 7–15 mm, glabrous, minutely hirtellous, or stipitate-glandular, throat closing, proximalmost lobe pair upcurving, distalmost 3–5 mm, prominently protruding.

greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect.

Capsules

included, 4–5 mm.

included, 5–8.5 mm.

Anthers

included, glabrous.

= 32.

Erythranthe caespitosa

Erythranthe washingtonensis

Phenology

Flowering Jul–Sep.

Flowering May–Sep.

Habitat

Alpine meadows and slopes, stream banks, wet rocks in streams, wet crevices, talus.

Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps.

Elevation

1100–2000(–2300) m. (3600–6600(–7500) ft.)

700–1300 m. (2300–4300 ft.)

Distribution

WA; BC

[WildflowerSearch map]

Discussion

Erythranthe caespitosa is endemic to northwestern and central Washington (Cascade and Olympic mountains) and adjacent British Columbia (Cascades, Selkirk Mountains and Chilliwack Valley, Coast Mountains). The plants have consistently small leaves with subentire margins, and the stems are consistently procumbent to decumbent-ascending, usually forming matted colonies. Erythranthe caespitosa and E. tilingii appear to be sympatric in counties of northwestern Washington, but this needs to be verified in the field.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 409.

FNA vol. 17, p. 396.

Parent taxa

Phrymaceae > Erythranthe

Sibling taxa

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. willisii

Synonyms

Mimulus scouleri var. caespitosus, M. caespitosus, M. tilingii var. caespitosus

Mimulus washingtonensis

Name authority

(Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012)

(Gandoger) G. L. Nesom: Phytoneuron 2012-39: 39. (2012) — (as washingtoniensis)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Erythranthe caespitosa

Erythranthe washingtonensis

Erythranthe caespitosa

Erythranthe washingtonensis