Erythranthe caespitosa |
Erythranthe scouleri |
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large mountain monkey-flower, mountain monkeyflower, Olympic monkeyflower, subalpine monkeyflower |
Columbia River monkey-flower, Scouler's monkey-flower |
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Habit | Perennials, rhizomatous, rooting at proximal nodes, sometimes producing creeping, small-leaved runners, forming matted colonies, rhizomes filiform. | Perennials, rhizomatous, producing long, sparsely leafy runners from basal nodes. |
Stems | procumbent or decumbent to decumbent-ascending, delicate, usually in masses, terete or flattish, branched, 3–10 cm, glabrous, minutely hirtellous, or stipitate-glandular. |
erect, simple or few-branched, 15–80 cm, glabrous. |
Leaves | basal and cauline; petiole: proximals 2–5 mm, distals 0 mm; blade often purple beneath, palmately 3-veined, orbicular to narrowly elliptic or ovate, proximals usually sublyrate, 3–12 mm, becoming larger distally, base cuneate to a short petiole, margins entire, mucronulate, or barely denticulate, apex obtuse, surfaces sparsely to moderately puberulent, hairs minute, stipitate-glandular. |
cauline, basal usually not persistent; petiole: proximals and medial 10–25 mm (gradually merging into blade), distals 0 mm; blade palmately (3–)5–7-veined to subpinnate, (proximal to medial) oblong-elliptic to oblong-lanceolate, 25–60 × 8–18 mm, usually 3–4 times longer than wide, base attenuate, margins evenly, shallowly dentate or crenate to mucronate or mucronulate, teeth 10–20 per side, sometimes more deeply toothed at base, apex obtuse to acute, surfaces glabrous. |
Flowers | herkogamous, 1–3, from distal nodes, commonly solitary. |
herkogamous, (1 or)2–8, from distal nodes. |
Styles | minutely hirtellous. |
minutely, prominently hirsutulous to villosulous. |
Corollas | yellow, dark red-spotted, bilaterally symmetric, bilabiate; tube-throat broadly funnelform to cylindric-funnelform, 15–18 mm, exserted beyond calyx margin; abaxial limb with deflexed-spreading lobes, adaxial with ascending lobes, palate partially closed. |
yellow, without red markings, bilaterally symmetric, bilabiate; tube-throat funnelform, 20–24 mm, exserted 10–15 mm beyond calyx margin; limb expanded 22–30 mm. |
Fruiting pedicels | 10–30(–40) mm, sparsely to moderately villous, hairs short, gland-tipped, sometimes hirtellous. |
20–25 mm, glabrous. |
Fruiting calyces | broadly campanulate, inflated, sagittally compressed, 7–15 mm, glabrous, minutely hirtellous, or stipitate-glandular, throat closing, proximalmost lobe pair upcurving, distalmost 3–5 mm, prominently protruding. |
ovoid, inflated, sagittally compressed, 13–14 mm, glabrous, throat closing. |
Capsules | included, 4–5 mm. |
unknown. |
Anthers | included, glabrous. |
included, glabrous. |
Erythranthe caespitosa |
Erythranthe scouleri |
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Phenology | Flowering Jul–Sep. | Flowering May–Aug. |
Habitat | Alpine meadows and slopes, stream banks, wet rocks in streams, wet crevices, talus. | Salt marshes, river banks. |
Elevation | 1100–2000(–2300) m. (3600–6600(–7500) ft.) | 0–20 m. (0–100 ft.) |
Distribution |
WA; BC
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OR; WA |
Discussion | Erythranthe caespitosa is endemic to northwestern and central Washington (Cascade and Olympic mountains) and adjacent British Columbia (Cascades, Selkirk Mountains and Chilliwack Valley, Coast Mountains). The plants have consistently small leaves with subentire margins, and the stems are consistently procumbent to decumbent-ascending, usually forming matted colonies. Erythranthe caespitosa and E. tilingii appear to be sympatric in counties of northwestern Washington, but this needs to be verified in the field. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe scouleri is distinctive in its oblong-elliptic leaves with long-tapering bases and closely toothed margins, completely glabrous vestiture, long, leafy runners from basal cauline nodes, large corollas with broad limbs, and prominently hairy styles. Several features suggest a close relationship to E. decora, particularly its very large corollas, hairy styles, closely toothed leaf margins, tall, simple, and erect stems, numerous runners, and its geographic range. All collections apparently have been made near the mouth of the Columbia River in Clatsop and Columbia counties, Oregon (G. L. Nesom 2013d). Recent observations (Alexander John Wright, pers. comm.; photos http://www.inaturalist.org/observations/3537008) indicate that it also occurs in Wahkiakum County, Washington, in the Julia Butler Hansen Refuge. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 409. | FNA vol. 17, p. 415. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus scouleri var. caespitosus, M. caespitosus, M. tilingii var. caespitosus | Mimulus scouleri, M. guttatus subsp. scouleri |
Name authority | (Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012) | (Hooker) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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