FNA: Erythranthe caespitosa vs. Erythranthe brachystylis

	Erythranthe caespitosa	Erythranthe brachystylis
	large mountain monkey-flower, mountain monkeyflower, Olympic monkeyflower, subalpine monkeyflower	short-pedicel monkeyflower
Habit	Perennials, rhizomatous, rooting at proximal nodes, sometimes producing creeping, small-leaved runners, forming matted colonies, rhizomes filiform.	Annuals, fibrous-rooted, sometimes taprooted, apparently sometimes producing thin runners from basal nodes.
Stems	procumbent or decumbent to decumbent-ascending, delicate, usually in masses, terete or flattish, branched, 3–10 cm, glabrous, minutely hirtellous, or stipitate-glandular.	erect, simple or branched from proximal to medial nodes, 4-angled, filiform to slightly thickened, not distinctly fistulose, 6–22 cm, glabrous.
Leaves	basal and cauline; petiole: proximals 2–5 mm, distals 0 mm; blade often purple beneath, palmately 3-veined, orbicular to narrowly elliptic or ovate, proximals usually sublyrate, 3–12 mm, becoming larger distally, base cuneate to a short petiole, margins entire, mucronulate, or barely denticulate, apex obtuse, surfaces sparsely to moderately puberulent, hairs minute, stipitate-glandular.	basal and cauline; petiole: proximals 1–8 mm, distals 0 mm; blade palmately 3–5-veined, ovate to depressed-ovate or suborbicular, 10–40 × 6–25 mm, base truncate to subcordate, margins undulate, subentire, or weakly, irregularly dentate, apex rounded, surfaces: proximals and medials glabrous, distals villous, hairs thin-walled, flattened, vitreous and sharp-pointed, eglandular.
Flowers	herkogamous, 1–3, from distal nodes, commonly solitary.	plesiogamous, 4–10, from medial to distal nodes, cleistogamous.
Styles	minutely hirtellous.	glabrous.
Corollas	yellow, dark red-spotted, bilaterally symmetric, bilabiate; tube-throat broadly funnelform to cylindric-funnelform, 15–18 mm, exserted beyond calyx margin; abaxial limb with deflexed-spreading lobes, adaxial with ascending lobes, palate partially closed.	yellow, without red markings, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular; tube-throat narrowly cylindric, 7–9 mm, exserted 0–1 mm beyond calyx margin; limb expanded 3 mm.
Fruiting pedicels	10–30(–40) mm, sparsely to moderately villous, hairs short, gland-tipped, sometimes hirtellous.	5–10 mm in proximal axils, shorter than or equal to subtending leaves, 1–5 mm distally and flowers and fruits appearing sessile or subsessile, glabrous.
Fruiting calyces	broadly campanulate, inflated, sagittally compressed, 7–15 mm, glabrous, minutely hirtellous, or stipitate-glandular, throat closing, proximalmost lobe pair upcurving, distalmost 3–5 mm, prominently protruding.	red-tinged to sparsely purple-dotted or not, broadly elliptic-ovoid, inflated, sagittally compressed, 10–13 mm, minutely hirtellous, throat not or slightly closing.
Capsules	included, 4–5 mm.	included, stipitate, 4–5 mm.
Anthers	included, glabrous.	included, glabrous.

	Erythranthe caespitosa	Erythranthe brachystylis
Phenology	Flowering Jul–Sep.	Flowering Jun–Aug.
Habitat	Alpine meadows and slopes, stream banks, wet rocks in streams, wet crevices, talus.	Around springs, steep slopes.
Elevation	1100–2000(–2300) m. (3600–6600(–7500) ft.)	2100 m. (6900 ft.)
Distribution	WA; BC [WildflowerSearch map]	NV
Discussion	Erythranthe caespitosa is endemic to northwestern and central Washington (Cascade and Olympic mountains) and adjacent British Columbia (Cascades, Selkirk Mountains and Chilliwack Valley, Coast Mountains). The plants have consistently small leaves with subentire margins, and the stems are consistently procumbent to decumbent-ascending, usually forming matted colonies. Erythranthe caespitosa and E. tilingii appear to be sympatric in counties of northwestern Washington, but this needs to be verified in the field. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Erythranthe brachystylis is closely similar to E. arvensis. Plants of both are annual in duration and produce depressed-ovate leaves, the distal with vitreous-villous surfaces, and tiny corollas barely exserted from the calyx and probably cleistogamous. Vestiture of the distal leaves includes an admixture of eglandular sharp-pointed hairs, sometimes encountered in E. arvensis, though not typical, perhaps reflecting introgression from E. nasuta. The distinction of Erythranthe brachystylis from E. arvensis is primarily in its foreshortened pedicels and more inflated fruiting calyces. The fruiting calyces appear to be subsessile or on pedicels shorter or only equaling the subtending leaves. The difference is essentially qualitative but produces a distinctive aspect. Erythranthe brachystylis is known only from the type collection in Nye County, a region where E. arvensis has not been documented. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 409.	FNA vol. 17, p. 422.
Parent taxa	Phrymaceae > Erythranthe	Phrymaceae > Erythranthe
Sibling taxa	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms	Mimulus scouleri var. caespitosus, M. caespitosus, M. tilingii var. caespitosus	Mimulus brachystylis
Name authority	(Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012)	(Edwin) G. L. Nesom: Phytoneuron 2012-39: 43. (2012)
Web links	Local floras: BC, OR, WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe brachystylis

large mountain monkey-flower, mountain monkeyflower, Olympic monkeyflower, subalpine monkeyflower

short-pedicel monkeyflower

Habit

Perennials, rhizomatous, rooting at proximal nodes, sometimes producing creeping, small-leaved runners, forming matted colonies, rhizomes filiform.

Annuals, fibrous-rooted, sometimes taprooted, apparently sometimes producing thin runners from basal nodes.

Stems

procumbent or decumbent to decumbent-ascending, delicate, usually in masses, terete or flattish, branched, 3–10 cm, glabrous, minutely hirtellous, or stipitate-glandular.

erect, simple or branched from proximal to medial nodes, 4-angled, filiform to slightly thickened, not distinctly fistulose, 6–22 cm, glabrous.

Leaves

basal and cauline;

petiole: proximals 2–5 mm, distals 0 mm;

blade often purple beneath, palmately 3-veined, orbicular to narrowly elliptic or ovate, proximals usually sublyrate, 3–12 mm, becoming larger distally, base cuneate to a short petiole, margins entire, mucronulate, or barely denticulate, apex obtuse, surfaces sparsely to moderately puberulent, hairs minute, stipitate-glandular.

basal and cauline;

petiole: proximals 1–8 mm, distals 0 mm;

blade palmately 3–5-veined, ovate to depressed-ovate or suborbicular, 10–40 × 6–25 mm, base truncate to subcordate, margins undulate, subentire, or weakly, irregularly dentate, apex rounded, surfaces: proximals and medials glabrous, distals villous, hairs thin-walled, flattened, vitreous and sharp-pointed, eglandular.

Flowers

herkogamous, 1–3, from distal nodes, commonly solitary.

plesiogamous, 4–10, from medial to distal nodes, cleistogamous.

Styles

minutely hirtellous.

glabrous.

Corollas

yellow, dark red-spotted, bilaterally symmetric, bilabiate;

tube-throat broadly funnelform to cylindric-funnelform, 15–18 mm, exserted beyond calyx margin;

abaxial limb with deflexed-spreading lobes, adaxial with ascending lobes, palate partially closed.

yellow, without red markings, weakly bilaterally or nearly radially symmetric, weakly bilabiate or nearly regular;

tube-throat narrowly cylindric, 7–9 mm, exserted 0–1 mm beyond calyx margin;

limb expanded 3 mm.

Fruiting pedicels

10–30(–40) mm, sparsely to moderately villous, hairs short, gland-tipped, sometimes hirtellous.

5–10 mm in proximal axils, shorter than or equal to subtending leaves, 1–5 mm distally and flowers and fruits appearing sessile or subsessile, glabrous.

Fruiting calyces

broadly campanulate, inflated, sagittally compressed, 7–15 mm, glabrous, minutely hirtellous, or stipitate-glandular, throat closing, proximalmost lobe pair upcurving, distalmost 3–5 mm, prominently protruding.

red-tinged to sparsely purple-dotted or not, broadly elliptic-ovoid, inflated, sagittally compressed, 10–13 mm, minutely hirtellous, throat not or slightly closing.

Capsules

included, 4–5 mm.

included, stipitate, 4–5 mm.

Anthers

included, glabrous.

Erythranthe caespitosa

Erythranthe brachystylis

Phenology

Flowering Jul–Sep.

Flowering Jun–Aug.

Habitat

Alpine meadows and slopes, stream banks, wet rocks in streams, wet crevices, talus.

Around springs, steep slopes.

Elevation

1100–2000(–2300) m. (3600–6600(–7500) ft.)

2100 m. (6900 ft.)

Distribution

WA; BC

[WildflowerSearch map]

Discussion

Erythranthe caespitosa is endemic to northwestern and central Washington (Cascade and Olympic mountains) and adjacent British Columbia (Cascades, Selkirk Mountains and Chilliwack Valley, Coast Mountains). The plants have consistently small leaves with subentire margins, and the stems are consistently procumbent to decumbent-ascending, usually forming matted colonies. Erythranthe caespitosa and E. tilingii appear to be sympatric in counties of northwestern Washington, but this needs to be verified in the field.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erythranthe brachystylis is closely similar to E. arvensis. Plants of both are annual in duration and produce depressed-ovate leaves, the distal with vitreous-villous surfaces, and tiny corollas barely exserted from the calyx and probably cleistogamous. Vestiture of the distal leaves includes an admixture of eglandular sharp-pointed hairs, sometimes encountered in E. arvensis, though not typical, perhaps reflecting introgression from E. nasuta.

The distinction of Erythranthe brachystylis from E. arvensis is primarily in its foreshortened pedicels and more inflated fruiting calyces. The fruiting calyces appear to be subsessile or on pedicels shorter or only equaling the subtending leaves. The difference is essentially qualitative but produces a distinctive aspect.

Erythranthe brachystylis is known only from the type collection in Nye County, a region where E. arvensis has not been documented.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 409.

FNA vol. 17, p. 422.

Parent taxa

Phrymaceae > Erythranthe

Sibling taxa

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

Synonyms

Mimulus scouleri var. caespitosus, M. caespitosus, M. tilingii var. caespitosus

Mimulus brachystylis

Name authority

(Greene) G. L. Nesom: Phytoneuron 2012-39: 43. (2012)

(Edwin) G. L. Nesom: Phytoneuron 2012-39: 43. (2012)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Erythranthe caespitosa

Erythranthe brachystylis

Erythranthe caespitosa

Erythranthe brachystylis