FNA: Erythranthe breweri vs. Erythranthe percaulis

	Erythranthe breweri	Erythranthe percaulis
	Brewer's monkey-flower	serpentine canyon monkeyflower
Habit	Annuals, taprooted.	Annuals, fibrous-rooted.
Stems	erect, simple or branched from basal nodes, 2–20 cm, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped.	erect or slightly ascending from base, often purplish, simple or few-branched from basal nodes, 7–28 cm, glabrous.
Leaves	cauline, basal not persistent; petiole 1–4 mm; blade 1-veined or palmately 3-veined (in broader ones), linear-oblanceolate to linear-lanceolate or linear, (5–)10–15(–20) × 1–2(–5) mm, base attenuate, margins entire or remotely mucronulate distally, apex acute, surfaces densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped.	basal and cauline; petiole: basal and proximalmost cauline 5–10 mm or cauline 0 mm; blade often spreading at right angles to stem, purple, palmately 3–5-veined, narrowly ovate, rhombic-elliptic, ovate to lanceolate, or elliptic-lanceolate to ovate or oblong-ovate, 7–10 mm, cauline even-sized or slightly smaller distally, 4–10 mm, base truncate to attenuate, margins entire or proximals shallowly sinuate, serrations 2–4, shallow, apex rounded, surfaces glabrous.
Flowers	herkogamous, 1–8, from medial to distal nodes.	herkogamous, 8–12, usually on distal 2/3 of stem, not clustered, chasmogamous.
Styles	glabrous.	glabrous.
Corollas	pink to red, rose red, red-purple, or lavender, throat usually light purple to lavender (similar in color to rest of corolla), palate ridges yellow, abaxial limb with deep pink markings, bilaterally symmetric, strongly bilabiate; tube-throat cylindric-funnelform, 3–7 mm, exserted 0–1 mm beyond calyx margin; limb expanded 2–3 mm, lobes notched or entire.	yellow, throat floor with a few red dots, proximal middle lip base with a larger red splotch, bilaterally symmetric, bilabiate; tube-throat cylindric-funnelform, 4–6 mm, exserted 2–3 mm beyond calyx margin; limb expanded 4–5 mm, palate ridges yellow, densely hairy.
Fruiting pedicels	4–12 mm, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped.	15–35 mm, glabrous.
Fruiting calyces	red-dotted, strongly angled, cylindric-campanulate, 4–7 mm, margins distinctly toothed or lobed, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped, ribs corky, lobes pronounced, spreading.	sharply wing-angled, urceolate to urceolate-campanulate, inflated, sagittally compressed, (4–)5–6 mm, glabrous, throat closing.
Capsules	included, 3–7 mm.	included, 3 mm.
Anthers	included, glabrous.	included, glabrous.

	Erythranthe breweri	Erythranthe percaulis
Phenology	Flowering Jun–Aug(–Sep).	Flowering May–Jun.
Habitat	Seeps and springs, damp rocks, vernal creek beds, cliffs, granite outcrops, rocky ridges, gravelly areas, meadow edges, stream edges.	Soil pockets, crevices, and boulders on serpentine cliffs, slopes, and roadcuts.
Elevation	(700–)1300–3500 m. ((2300–)4300–11500 ft.)	2800 m. (9200 ft.)
Distribution	AZ; CA; ID; MT; NV; OR; UT; WA; WY; AB; BC [WildflowerSearch map]	CA
Discussion		Erythranthe percaulis was described from only the type collection from Serpentine Canyon of the Feather River in Plumas County, but the type locality has recently been relocated and the population determined to comprise many thousands of individuals (S. Schoenig 2016). Plants are characterized by their completely glabrous vestiture, terete and mostly simple stems, small leaves on relatively widely spaced nodes, small calyces, and small, yellow corollas with herkogamous arrangement of stigma and anthers. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 390.	FNA vol. 17, p. 418.
Parent taxa	Phrymaceae > Erythranthe	Phrymaceae > Erythranthe
Sibling taxa	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii	E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii
Synonyms	Eunanus breweri, Mimulus breweri, M. rubellus var. breweri
Name authority	(Greene) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 36. (2012)	G. L. Nesom: Phytoneuron 2013-70: 1, figs. 1–5. (2013)
Web links	Local floras: BC, CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe percaulis

Brewer's monkey-flower

serpentine canyon monkeyflower

Habit

Annuals, taprooted.

Annuals, fibrous-rooted.

Stems

erect, simple or branched from basal nodes, 2–20 cm, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped.

erect or slightly ascending from base, often purplish, simple or few-branched from basal nodes, 7–28 cm, glabrous.

Leaves

cauline, basal not persistent;

petiole 1–4 mm;

blade 1-veined or palmately 3-veined (in broader ones), linear-oblanceolate to linear-lanceolate or linear, (5–)10–15(–20) × 1–2(–5) mm, base attenuate, margins entire or remotely mucronulate distally, apex acute, surfaces densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped.

basal and cauline;

petiole: basal and proximalmost cauline 5–10 mm or cauline 0 mm;

blade often spreading at right angles to stem, purple, palmately 3–5-veined, narrowly ovate, rhombic-elliptic, ovate to lanceolate, or elliptic-lanceolate to ovate or oblong-ovate, 7–10 mm, cauline even-sized or slightly smaller distally, 4–10 mm, base truncate to attenuate, margins entire or proximals shallowly sinuate, serrations 2–4, shallow, apex rounded, surfaces glabrous.

Flowers

herkogamous, 1–8, from medial to distal nodes.

herkogamous, 8–12, usually on distal 2/3 of stem, not clustered, chasmogamous.

Styles

glabrous.

Corollas

pink to red, rose red, red-purple, or lavender, throat usually light purple to lavender (similar in color to rest of corolla), palate ridges yellow, abaxial limb with deep pink markings, bilaterally symmetric, strongly bilabiate;

tube-throat cylindric-funnelform, 3–7 mm, exserted 0–1 mm beyond calyx margin;

limb expanded 2–3 mm, lobes notched or entire.

yellow, throat floor with a few red dots, proximal middle lip base with a larger red splotch, bilaterally symmetric, bilabiate;

tube-throat cylindric-funnelform, 4–6 mm, exserted 2–3 mm beyond calyx margin;

limb expanded 4–5 mm, palate ridges yellow, densely hairy.

Fruiting pedicels

4–12 mm, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped.

15–35 mm, glabrous.

Fruiting calyces

red-dotted, strongly angled, cylindric-campanulate, 4–7 mm, margins distinctly toothed or lobed, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped, ribs corky, lobes pronounced, spreading.

sharply wing-angled, urceolate to urceolate-campanulate, inflated, sagittally compressed, (4–)5–6 mm, glabrous, throat closing.

Capsules

included, 3–7 mm.

included, 3 mm.

Anthers

included, glabrous.

Erythranthe breweri

Erythranthe percaulis

Phenology

Flowering Jun–Aug(–Sep).

Flowering May–Jun.

Habitat

Seeps and springs, damp rocks, vernal creek beds, cliffs, granite outcrops, rocky ridges, gravelly areas, meadow edges, stream edges.

Soil pockets, crevices, and boulders on serpentine cliffs, slopes, and roadcuts.

Elevation

(700–)1300–3500 m. ((2300–)4300–11500 ft.)

2800 m. (9200 ft.)

Distribution

AZ; CA; ID; MT; NV; OR; UT; WA; WY; AB; BC

[WildflowerSearch map]

Discussion

Erythranthe percaulis was described from only the type collection from Serpentine Canyon of the Feather River in Plumas County, but the type locality has recently been relocated and the population determined to comprise many thousands of individuals (S. Schoenig 2016). Plants are characterized by their completely glabrous vestiture, terete and mostly simple stems, small leaves on relatively widely spaced nodes, small calyces, and small, yellow corollas with herkogamous arrangement of stigma and anthers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 390.

FNA vol. 17, p. 418.

Parent taxa

Phrymaceae > Erythranthe

Sibling taxa

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. percaulis, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

E. acutidens, E. alsinoides, E. ampliata, E. androsacea, E. arenaria, E. arenicola, E. arvensis, E. barbata, E. bicolor, E. brachystylis, E. breviflora, E. breweri, E. caespitosa, E. calcicola, E. calciphila, E. cardinalis, E. carsonensis, E. charlestonensis, E. chinatiensis, E. cinnabarina, E. corallina, E. cordata, E. decora, E. dentata, E. diffusa, E. discolor, E. eastwoodiae, E. erubescens, E. exigua, E. filicaulis, E. filicifolia, E. floribunda, E. gemmipara, E. geniculata, E. geyeri, E. glaucescens, E. gracilipes, E. grandis, E. grayi, E. guttata, E. hallii, E. hardhamiae, E. hymenophylla, E. inamoena, E. inconspicua, E. inflatula, E. jungermannioides, E. laciniata, E. latidens, E. lewisii, E. linearifolia, E. marmorata, E. michiganensis, E. microphylla, E. minor, E. montioides, E. moschata, E. nasuta, E. norrisii, E. nudata, E. palmeri, E. pardalis, E. parishii, E. parvula, E. patula, E. primuloides, E. ptilota, E. pulsiferae, E. purpurea, E. regni, E. rhodopetra, E. rubella, E. scouleri, E. shevockii, E. sierrae, E. suksdorfii, E. taylorii, E. thermalis, E. tilingii, E. trinitiensis, E. unimaculata, E. utahensis, E. verbenacea, E. washingtonensis, E. willisii

Synonyms

Eunanus breweri, Mimulus breweri, M. rubellus var. breweri

Name authority

(Greene) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 36. (2012)

G. L. Nesom: Phytoneuron 2013-70: 1, figs. 1–5. (2013)

Web links

Local floras: BC, CA, OR, WA
Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria
WildflowerSearch
iNaturalist (observations)
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Flora of North America species comparison

Erythranthe breweri

Erythranthe percaulis

Erythranthe breweri

Erythranthe percaulis