Erythranthe breweri |
Erythranthe marmorata |
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Brewer's monkey-flower |
Stanislaus monkeyflower, Whipple's monkeyflower |
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Habit | Annuals, taprooted. | Annuals, taprooted. |
Stems | erect, simple or branched from basal nodes, 2–20 cm, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped. |
erect, simple or branched from base, 7–28 cm, usually moderately to densely villous-glandular, without eglandular hairs. |
Leaves | cauline, basal not persistent; petiole 1–4 mm; blade 1-veined or palmately 3-veined (in broader ones), linear-oblanceolate to linear-lanceolate or linear, (5–)10–15(–20) × 1–2(–5) mm, base attenuate, margins entire or remotely mucronulate distally, apex acute, surfaces densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped. |
usually cauline or basal persistent; petiole: proximals to medials 7–15 mm, distals 0 mm; blade palmately (3–)5-veined, ovate or broadly ovate to elliptic-ovate or depressed-ovate, (10–)15–30 × 6–15 mm, base truncate to shallowly cuneate, margins shallowly to coarsely dentate, apex acute, surfaces usually moderately to densely villous-glandular, without eglandular hairs. |
Flowers | herkogamous, 1–8, from medial to distal nodes. |
herkogamous, (1–)2–6, axillary from middle to distal nodes, chasmogamous. |
Styles | glabrous. |
glabrous or sparsely hirtellous. |
Corollas | pink to red, rose red, red-purple, or lavender, throat usually light purple to lavender (similar in color to rest of corolla), palate ridges yellow, abaxial limb with deep pink markings, bilaterally symmetric, strongly bilabiate; tube-throat cylindric-funnelform, 3–7 mm, exserted 0–1 mm beyond calyx margin; limb expanded 2–3 mm, lobes notched or entire. |
yellow, throat red-spotted, abaxial limb base with a large red splotch, bilaterally symmetric, bilabiate; tube-throat narrowly cylindric-funnelform, 10–12 mm, exserted 4–5 mm beyond calyx margin; limb abruptly expanded 14–20 mm. |
Fruiting pedicels | 4–12 mm, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped. |
15–45 mm, usually moderately to densely villous-glandular, without eglandular hairs. |
Fruiting calyces | red-dotted, strongly angled, cylindric-campanulate, 4–7 mm, margins distinctly toothed or lobed, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped, ribs corky, lobes pronounced, spreading. |
sharply nodding, usually densely purple-spotted, broadly campanulate, inflated, sagittally compressed, 9–12 mm, densely hirtellous, sometimes sparsely stipitate-glandular, densely villous at sinuses, throat closing, adaxial lobe ca. 2 times length of others. |
Capsules | included, 3–7 mm. |
included, 6–9 mm. |
Anthers | included, glabrous. |
included, glabrous. |
Erythranthe breweri |
Erythranthe marmorata |
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Phenology | Flowering Jun–Aug(–Sep). | Flowering Mar–May. |
Habitat | Seeps and springs, damp rocks, vernal creek beds, cliffs, granite outcrops, rocky ridges, gravelly areas, meadow edges, stream edges. | Habitat unknown, not over serpentine. |
Elevation | (700–)1300–3500 m. ((2300–)4300–11500 ft.) | 100–900 m. (300–3000 ft.) |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; WA; WY; AB; BC
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CA |
Discussion | Erythranthe marmorata is recognized by its erect, taprooted habit and annual duration, villous-glandular vestiture, ovate-petiolate leaves, flowers from middle to distal nodes, long, narrow corolla tube-throat abruptly flaring into a broad limb, abaxial middle corolla lobe with a large red splotch, and fruiting calyces dark-spotted and sharply nodding. The species is known from foothills in Calaveras, Stanislaus, and Tuolumne counties in the Stanislaus River drainage and from Amador County in the Mokelumne River drainage. A collection from Fresno County appears to be somewhat disjunct from the main range, and the plants are more densely villous than characteristic elsewhere, but their identification as E. marmorata otherwise seems secure. Erythranthe marmorata (previously identified as Mimulus whipplei) had been considered extremely rare or even perhaps extinct. See G. L. Nesom (2013d) for citations of recent collections. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 17, p. 390. | FNA vol. 17, p. 416. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Eunanus breweri, Mimulus breweri, M. rubellus var. breweri | Mimulus marmoratus, M. whipplei |
Name authority | (Greene) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 36. (2012) | (Greene) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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