Erythranthe breviflora |
Erythranthe scouleri |
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short-flower monkey-flower |
Columbia River monkey-flower, Scouler's monkey-flower |
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Habit | Annuals, shallowly fibrous-rooted. | Perennials, rhizomatous, producing long, sparsely leafy runners from basal nodes. |
Stems | ascending, geniculate at nodes, branched at proximal and medial nodes, 4–15 cm, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped, sometimes minutely hirtellous, hairs sharp-pointed, eglandular. |
erect, simple or few-branched, 15–80 cm, glabrous. |
Leaves | usually cauline, basal usually deciduous by flowering; petiole 1–3 mm; blade palmately 3-veined, narrowly ovate or narrowly lanceolate to elliptic or elliptic-lanceolate, largest 5–15 × 2–6 mm, relatively even-sized, or slightly reduced distally, base attenuate, margins entire, mucronulate, or denticulate, apex acute to obtuse, surfaces minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped, sometimes minutely hirtellous, hairs sharp-pointed, eglandular. |
cauline, basal usually not persistent; petiole: proximals and medial 10–25 mm (gradually merging into blade), distals 0 mm; blade palmately (3–)5–7-veined to subpinnate, (proximal to medial) oblong-elliptic to oblong-lanceolate, 25–60 × 8–18 mm, usually 3–4 times longer than wide, base attenuate, margins evenly, shallowly dentate or crenate to mucronate or mucronulate, teeth 10–20 per side, sometimes more deeply toothed at base, apex obtuse to acute, surfaces glabrous. |
Flowers | plesiogamous, 10–20, from medial to distal nodes. |
herkogamous, (1 or)2–8, from distal nodes. |
Styles | glabrous. |
minutely, prominently hirsutulous to villosulous. |
Corollas | yellow, red-spotted or striped, bilaterally symmetric, weakly bilabiate; tube-throat cylindric to narrowly funnelform, 3.5–5 mm, not exserted beyond calyx margin; limb barely widened, lobes broadly obovate, apex rounded. |
yellow, without red markings, bilaterally symmetric, bilabiate; tube-throat funnelform, 20–24 mm, exserted 10–15 mm beyond calyx margin; limb expanded 22–30 mm. |
Fruiting pedicels | straight, 5–11 mm, minutely stipitate-glandular, hairs 0.1–0.3 mm, gland-tipped, sometimes minutely hirtellous, hairs sharp-pointed, eglandular. |
20–25 mm, glabrous. |
Fruiting calyces | winged, plicate-angled, campanulate becoming ovoid-ellipsoid to campanulate, distinctly inflated, 5–6 mm, margins distinctly toothed or lobed, sparsely, minutely hirtellous, eglandular, sometimes sparsely sessile-glandular, lobes pronounced, erect. |
ovoid, inflated, sagittally compressed, 13–14 mm, glabrous, throat closing. |
Capsules | included, 4–6 mm. |
unknown. |
Anthers | included, glabrous. |
included, glabrous. |
Erythranthe breviflora |
Erythranthe scouleri |
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Phenology | Flowering May–Jul. | Flowering May–Aug. |
Habitat | Stream and lake sides, gravel bars, springs, moist slopes, damp swales between dunes, along trails. | Salt marshes, river banks. |
Elevation | 700–2300 m. (2300–7500 ft.) | 0–20 m. (0–100 ft.) |
Distribution |
CA; ID; MT; NV; OR; WA; BC
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OR; WA |
Discussion | Erythranthe scouleri is distinctive in its oblong-elliptic leaves with long-tapering bases and closely toothed margins, completely glabrous vestiture, long, leafy runners from basal cauline nodes, large corollas with broad limbs, and prominently hairy styles. Several features suggest a close relationship to E. decora, particularly its very large corollas, hairy styles, closely toothed leaf margins, tall, simple, and erect stems, numerous runners, and its geographic range. All collections apparently have been made near the mouth of the Columbia River in Clatsop and Columbia counties, Oregon (G. L. Nesom 2013d). Recent observations (Alexander John Wright, pers. comm.; photos http://www.inaturalist.org/observations/3537008) indicate that it also occurs in Wahkiakum County, Washington, in the Julia Butler Hansen Refuge. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 17, p. 399. | FNA vol. 17, p. 415. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus breviflorus | Mimulus scouleri, M. guttatus subsp. scouleri |
Name authority | (Piper) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) | (Hooker) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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