Erythranthe bicolor |
Erythranthe geniculata |
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yellow and white monkeyflower |
bent-stem monkeyflower, Dudley's monkeyflower |
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Habit | Annuals, taprooted. | Annuals, fibrous-rooted or filiform-taprooted. |
Stems | erect, simple or branched from basal nodes, 4–30 cm, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped. |
ascending to decumbent or prostrate, geniculate at nodes, simple or diffusely branched, 5–60 cm, moderately villous, hairs 0.8–2 mm, multicellular, eglandular and also 0.1–0.3 mm, stipitate-glandular. |
Leaves | cauline, basal not persistent; petiole 0 mm or short-attenuate; blade 1-veined or palmately 3-veined (in broader ones), narrowly elliptic to linear-lanceolate or linear-oblanceolate, 10–30 × 2–6 mm at mid stem, base attenuate, margins entire or remotely shallowly dentate to mucronulate, teeth 1–4 per side, apex acute, surfaces densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped. |
basal and cauline, basal usually deciduous by flowering; petiole 2–10(–35) mm; blade pinnately to subpinnately veined, broadly ovate or elliptic-ovate to triangular, 8–35 × 5–30 mm, base cuneate to rounded or subcordate, margins serrate or dentate, teeth 3–10 per side, apex acute to obtuse or rounded, surfaces moderately villous, hairs 0.8–2 mm, multicellular, eglandular, and 0.1–0.3 mm, stipitate-glandular. |
Flowers | herkogamous, 2–8, from medial to distal nodes. |
herkogamous, (1–)6–20, from all or medial to distal nodes. |
Styles | minutely glandular. |
glabrous. |
Corollas | yellow, adaxial lip white, sometimes yellow, throat and abaxial limb red-spotted, bilaterally symmetric, weakly bilabiate; tube-throat funnelform, 8–12 mm, exserted beyond calyx margin; limb broadly expanded, lobes 2-fid, palate villous. |
yellow, without white patches, throat red-spotted, spots concentrated or becoming coalescent into a somewhat discrete splotch at base of each of 3 abaxial lobes and sometimes 2 adaxial, bilaterally symmetric, ± bilabiate; tube-throat cylindric, 9–12 mm, exserted beyond calyx margin; limb expanded 10–18 mm diam. |
Fruiting pedicels | (5–)7–20(–30) mm, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped. |
12–26(–55) mm, moderately villous, hairs 0.8–2 mm, multicellular, eglandular and also 0.1–0.3 mm, stipitate-glandular. |
Fruiting calyces | red-dotted, strongly angled, cylindric-campanulate, (5–)7–12 mm, margins distinctly toothed or lobed, densely glandular-puberulent, hairs 0.05–0.1 mm, gland-tipped, ribs corky, lobes pronounced, spreading. |
red-spotted, campanulate-cylindric, weakly inflated, (5–)6–8 mm, margins distinctly toothed or lobed, sparsely to moderately villous-glandular, ribs shallowly wing-angled, lobes pronounced, erect to spreading or spreading-recurving. |
Capsules | included, 4–6 mm. |
included, 4–6(–7) mm. |
Anthers | included, margins ciliate, glabrous. |
included, glabrous. |
2n | = 32. |
|
Erythranthe bicolor |
Erythranthe geniculata |
|
Phenology | Flowering Apr–Jun. | Flowering (Mar–)Apr–Jul. |
Habitat | Moist banks, serpentine and granite outcrops, seepage areas, volcanic mudflows, open red clay exposures. | Granite crevices, canyon slopes, talus, crevices in volcanic outcrops, edges of boulders, roadsides, damp sandy soils, sandy water edges, gravelly soils and creek bottoms. |
Elevation | (100–)400–1700(–2500) m. ((300–)1300–5600(–8200) ft.) | 200–900(–1200) m. (700–3000(–3900) ft.) |
Distribution |
CA
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CA
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Discussion | Erythranthe bicolor ranges from Shasta and Trinity counties south to Tulare County; the identity of apparently disjunct populations in San Bernardino County needs to be reexamined. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe geniculata is known from an apparently disjunct cluster of populations in Butte, Sutter, and Yuba counties and then from Tuolumne and Stanislaus counties south to Kern County. Erythranthe geniculata, compared to E. floribunda, has larger, chasmogamous, and allogamous flowers. The anther pairs of E. geniculata are at different levels, and the stigma is slightly above the adaxial anther pair; in E. floribunda both anther pairs and the stigma are at the same level. Erythranthe arenaria, E. geniculata, and E. norrisii constitute a group of apparently closely related species within sect. Mimulosma endemic along the Sierra Nevada. All have ovate-petiolate leaves (only the basal ones are sometimes ovate in E. arenaria) with pinnate to subpinnate venation. The more widespread E. floribunda, which is part of the above group, also is similar, but all three endemics have larger corollas with the tube exserted at greater length beyond the calyx margin. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 390. | FNA vol. 17, p. 404. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus bicolor | Mimulus geniculatus, M. dudleyi, M. floribundus var. geniculatus |
Name authority | (Hartweg ex Bentham) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 36. (2012) | (Greene) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) |
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