Erythranthe arenaria |
Erythranthe utahensis |
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sand-loving monkeyflower |
Utah monkeyflower |
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Habit | Annuals, fibrous-rooted or filiform-taprooted. | Perennials, rhizomatous, rooting at nodes, rhizomes filiform. |
Stems | erect to ascending, straight or geniculate at nodes, simple or branched, 5–20 cm, villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
erect, sometimes decumbent-ascending proximally, simple or few-branched, 20–50 cm, glabrous or sparsely stipitate-glandular in inflorescence. |
Leaves | basal and cauline; petiole 0 mm or proximals 1–3(–5) mm; blade 1-veined or palmately 3-veined, elliptic to narrowly elliptic, ovate-elliptic, or ovate-lanceolate, 5–12(–17) × 3–7 mm, base rounded to cuneate-attenuate, margins entire or sparsely dentate to serrate, apex acuminate to acute or obtuse, surfaces villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
basal and cauline, even-sized or largest near mid stem; petiole 0 mm or proximalmost 2–10 mm; blade palmately 3–5-veined, orbicular or suborbicular to broadly elliptic, broadly ovate, or depressed-ovate, 20–40(–75) × 12–35(–40) mm, base usually truncate to broadly cuneate, margins entire or subentire to mucronulate, shallowly dentate, or denticulate, apex rounded, surfaces glabrous or glabrate to sparsely stipitate-glandular and sparsely pilose, hairs thin-walled, abaxial often glaucous. |
Flowers | herkogamous, 1–22, from proximal to distal nodes. |
herkogamous, 6–16, from proximal to distal nodes, in a loose raceme, distal bracts becoming much reduced. |
Styles | glabrous. |
hirtellous. |
Corollas | yellow, abaxial limb red-dotted, bilaterally symmetric, weakly bilabiate; tube-throat funnelform, 9–12(–14) mm, exserted beyond calyx margin; lobes broadly obovate, apex rounded. |
yellow, abaxial limb prominently darker yellow, sparsely purple-spotted, bilaterally symmetric, weakly bilabiate; tube-throat narrowly funnelform to broadly cylindric, 10–15 mm, exserted 5–8 mm beyond calyx margin; limb expanded 12–20 mm. |
Fruiting pedicels | divergent-arcuate, 10–23 mm, villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
(25–)40–75 mm, stipitate-glandular to short villous, hairs gland-tipped. |
Fruiting calyces | usually red-dotted, narrowly campanulate, not or weakly inflated, 5–7(–9) mm, margins distinctly toothed or lobed, villous-glandular, ribs angled, lobes pronounced, erect. |
broadly ovate-cylindric, inflated, sagittally compressed, (10–)11–17(–20) mm, stipitate-glandular or minutely hirtellous or a mixture, hairs sometimes also longer, thin-walled, eglandular or glandular, throat not closing, adaxial lobe slightly longer, triangular-blunt. |
Capsules | included, 4–7 mm. |
included, 5–7 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 32. |
= 28, 30. |
Erythranthe arenaria |
Erythranthe utahensis |
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Phenology | Flowering May–Sep. | Flowering (Feb–)May–Aug(–Oct). |
Habitat | Sandy flats, bars, gullies, washes, trails, roadcuts, seasonal creek beds and drainages, rocky slopes, seepy loam, ditches, lake edges, meadows, openings in pine-fir and pine-oak woodlands. | Drainage ditches, springs, seeps, wet meadows, margins of ponds and small streams, marshy areas. |
Elevation | (100–)500–2600(–2800) m. ((300–)1600–8500(–9200) ft.) | 1400–2500 m. (4600–8200 ft.) |
Distribution |
CA
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CA; CO; NV; UT |
Discussion | Erythranthe arenaria is known from a cluster of six counties of the central Sierra Nevada: Fresno, Madera, Mariposa, Merced, Tulare, and Tuolumne. Most plants of Erythranthe arenaria have relatively even-sized cauline leaves, all sessile to proximally subsessile. Plants in the Yosemite area with persistent basal leaves that are short-petiolate, ovate with a cuneate base, and relatively larger than the more distal cauline ones, and possibly related to E. arenaria, have been named M. floribundus var. subulatus. These might be construed as showing the influence of E. geniculata, but that species occurs only at the lower range of elevation of E. arenaria, while plants referable to Mimulus floribundus var. subulatus occur at least to 2300 m and also have the erect habit characteristic of E. arenaria. These variants should be investigated, especially in the Yosemite area where they appear to be relatively common, with the possibility that they indeed represent a distinct entity. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe utahensis is characterized by its erect stems, prolifically produced filiform rhizomes, basal leaves short-petiolate to subsessile and cauline sessile, blades suborbicular to broadly ovate or depressed ovate with thin-walled villous-glandular hairs on both surfaces, margins subentire, proximal pedicels elongating to 75 mm, and calyces open at maturity. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 403. | FNA vol. 17, p. 410. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus arenarius, M. floribundus var. subulatus, M. multiflorus, M. subulatus, M. trisulcatus | Mimulus glabratus subsp. utahensis |
Name authority | (A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) | (Pennell) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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