Erythranthe arenaria |
Erythranthe unimaculata |
|
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sand-loving monkeyflower |
green-palate monkeyflower |
|
Habit | Annuals, fibrous-rooted or filiform-taprooted. | Annuals, shallowly fibrous-rooted or slender-taprooted, sometimes rooting at proximal nodes. |
Stems | erect to ascending, straight or geniculate at nodes, simple or branched, 5–20 cm, villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
erect or basally ascending-erect, simple or few-branched, becoming fistulose in larger plants, (2–)10–30(–100) cm, delicately short glandular-villous to stipitate-glandular, often glabrous below inflorescence. |
Leaves | basal and cauline; petiole 0 mm or proximals 1–3(–5) mm; blade 1-veined or palmately 3-veined, elliptic to narrowly elliptic, ovate-elliptic, or ovate-lanceolate, 5–12(–17) × 3–7 mm, base rounded to cuneate-attenuate, margins entire or sparsely dentate to serrate, apex acuminate to acute or obtuse, surfaces villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
usually cauline, basal sometimes persistent, distal connate-perfoliate, often bractlike; petiole: proximals 4–10(–15) mm, distals 0 mm; blade: proximals sometimes subpinnately veined, usually with (1 or)2 pairs arising from midvein above base, becoming palmately veined distally, ovate-lanceolate to ovate or broadly ovate-elliptic, mid cauline 12–40(–50) × 10–25(–45) mm, base rounded to truncate or cuneate, margins shallowly dentate-serrate to serrate, teeth 7–12 per side, apex rounded to obtuse, surfaces glabrous, sometimes with sharp-pointed hirtellous, vitreous-flattened, or gland-tipped hairs. |
Flowers | herkogamous, 1–22, from proximal to distal nodes. |
herkogamous, (1–)3–14, usually from mid stem and distally, chasmogamous. |
Styles | glabrous. |
hirtellous. |
Corollas | yellow, abaxial limb red-dotted, bilaterally symmetric, weakly bilabiate; tube-throat funnelform, 9–12(–14) mm, exserted beyond calyx margin; lobes broadly obovate, apex rounded. |
pale yellow, palate and abaxial throat dark yellow, drying blue-green, red-spotted, bilaterally symmetric, bilabiate; tube-throat funnelform to subfunnelform, (7–)9–14 mm, exserted 3–4 mm beyond calyx margin; limb broadly expanded (8–17 mm pressed), palate densely bearded. |
Fruiting pedicels | divergent-arcuate, 10–23 mm, villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
10–40 mm, delicately short glandular-villous to stipitate-glandular, often glabrous below inflorescence. |
Fruiting calyces | usually red-dotted, narrowly campanulate, not or weakly inflated, 5–7(–9) mm, margins distinctly toothed or lobed, villous-glandular, ribs angled, lobes pronounced, erect. |
nodding 30–90º, broadly campanulate, inflated, sagittally compressed, 9–13(–15) mm, sparsely glandular-villous to stipitate-glandular, throat closing, adaxial lobe distinctly longer. |
Capsules | included, 4–7 mm. |
included, 5–8 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 32. |
= 28. |
Erythranthe arenaria |
Erythranthe unimaculata |
|
Phenology | Flowering May–Sep. | Flowering Jan–Jun. |
Habitat | Sandy flats, bars, gullies, washes, trails, roadcuts, seasonal creek beds and drainages, rocky slopes, seepy loam, ditches, lake edges, meadows, openings in pine-fir and pine-oak woodlands. | Stream and canal sides, pool edges, canyon bottoms, sand, gravel, and mud, riparian habitats, pine-oak forests. |
Elevation | (100–)500–2600(–2800) m. ((300–)1600–8500(–9200) ft.) | 200–2000 m. (700–6600 ft.) |
Distribution |
CA
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AZ; NM; Mexico (Chihuahua, Sonora) |
Discussion | Erythranthe arenaria is known from a cluster of six counties of the central Sierra Nevada: Fresno, Madera, Mariposa, Merced, Tulare, and Tuolumne. Most plants of Erythranthe arenaria have relatively even-sized cauline leaves, all sessile to proximally subsessile. Plants in the Yosemite area with persistent basal leaves that are short-petiolate, ovate with a cuneate base, and relatively larger than the more distal cauline ones, and possibly related to E. arenaria, have been named M. floribundus var. subulatus. These might be construed as showing the influence of E. geniculata, but that species occurs only at the lower range of elevation of E. arenaria, while plants referable to Mimulus floribundus var. subulatus occur at least to 2300 m and also have the erect habit characteristic of E. arenaria. These variants should be investigated, especially in the Yosemite area where they appear to be relatively common, with the possibility that they indeed represent a distinct entity. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe unimaculata is recognized by its annual duration (fibrous-rooted, without stolons or rhizomes), delicate-glandular vestiture, mostly sessile to subsessile, often widely spaced leaves, closed fruiting calyces, and relatively large, pale yellow to nearly white corollas with a dark yellow palate that commonly dries blue-green. The breeding system is allogamous. Plants commonly are single-stemmed and usually erect but sometimes produce decumbent-ascending stems branching from the base, these sometimes rooting at proximal nodes. Erythranthe unimaculata is known from Cochise, Gila, Pima, and Santa Cruz counties, Arizona, and from Doña Ana County, New Mexico. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 403. | FNA vol. 17, p. 413. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus arenarius, M. floribundus var. subulatus, M. multiflorus, M. subulatus, M. trisulcatus | Mimulus unimaculatus |
Name authority | (A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) | (Pennell) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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