Erythranthe arenaria |
Erythranthe percaulis |
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sand-loving monkeyflower |
serpentine canyon monkeyflower |
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Habit | Annuals, fibrous-rooted or filiform-taprooted. | Annuals, fibrous-rooted. |
Stems | erect to ascending, straight or geniculate at nodes, simple or branched, 5–20 cm, villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
erect or slightly ascending from base, often purplish, simple or few-branched from basal nodes, 7–28 cm, glabrous. |
Leaves | basal and cauline; petiole 0 mm or proximals 1–3(–5) mm; blade 1-veined or palmately 3-veined, elliptic to narrowly elliptic, ovate-elliptic, or ovate-lanceolate, 5–12(–17) × 3–7 mm, base rounded to cuneate-attenuate, margins entire or sparsely dentate to serrate, apex acuminate to acute or obtuse, surfaces villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
basal and cauline; petiole: basal and proximalmost cauline 5–10 mm or cauline 0 mm; blade often spreading at right angles to stem, purple, palmately 3–5-veined, narrowly ovate, rhombic-elliptic, ovate to lanceolate, or elliptic-lanceolate to ovate or oblong-ovate, 7–10 mm, cauline even-sized or slightly smaller distally, 4–10 mm, base truncate to attenuate, margins entire or proximals shallowly sinuate, serrations 2–4, shallow, apex rounded, surfaces glabrous. |
Flowers | herkogamous, 1–22, from proximal to distal nodes. |
herkogamous, 8–12, usually on distal 2/3 of stem, not clustered, chasmogamous. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, abaxial limb red-dotted, bilaterally symmetric, weakly bilabiate; tube-throat funnelform, 9–12(–14) mm, exserted beyond calyx margin; lobes broadly obovate, apex rounded. |
yellow, throat floor with a few red dots, proximal middle lip base with a larger red splotch, bilaterally symmetric, bilabiate; tube-throat cylindric-funnelform, 4–6 mm, exserted 2–3 mm beyond calyx margin; limb expanded 4–5 mm, palate ridges yellow, densely hairy. |
Fruiting pedicels | divergent-arcuate, 10–23 mm, villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
15–35 mm, glabrous. |
Fruiting calyces | usually red-dotted, narrowly campanulate, not or weakly inflated, 5–7(–9) mm, margins distinctly toothed or lobed, villous-glandular, ribs angled, lobes pronounced, erect. |
sharply wing-angled, urceolate to urceolate-campanulate, inflated, sagittally compressed, (4–)5–6 mm, glabrous, throat closing. |
Capsules | included, 4–7 mm. |
included, 3 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 32. |
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Erythranthe arenaria |
Erythranthe percaulis |
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Phenology | Flowering May–Sep. | Flowering May–Jun. |
Habitat | Sandy flats, bars, gullies, washes, trails, roadcuts, seasonal creek beds and drainages, rocky slopes, seepy loam, ditches, lake edges, meadows, openings in pine-fir and pine-oak woodlands. | Soil pockets, crevices, and boulders on serpentine cliffs, slopes, and roadcuts. |
Elevation | (100–)500–2600(–2800) m. ((300–)1600–8500(–9200) ft.) | 2800 m. (9200 ft.) |
Distribution |
CA
|
CA |
Discussion | Erythranthe arenaria is known from a cluster of six counties of the central Sierra Nevada: Fresno, Madera, Mariposa, Merced, Tulare, and Tuolumne. Most plants of Erythranthe arenaria have relatively even-sized cauline leaves, all sessile to proximally subsessile. Plants in the Yosemite area with persistent basal leaves that are short-petiolate, ovate with a cuneate base, and relatively larger than the more distal cauline ones, and possibly related to E. arenaria, have been named M. floribundus var. subulatus. These might be construed as showing the influence of E. geniculata, but that species occurs only at the lower range of elevation of E. arenaria, while plants referable to Mimulus floribundus var. subulatus occur at least to 2300 m and also have the erect habit characteristic of E. arenaria. These variants should be investigated, especially in the Yosemite area where they appear to be relatively common, with the possibility that they indeed represent a distinct entity. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe percaulis was described from only the type collection from Serpentine Canyon of the Feather River in Plumas County, but the type locality has recently been relocated and the population determined to comprise many thousands of individuals (S. Schoenig 2016). Plants are characterized by their completely glabrous vestiture, terete and mostly simple stems, small leaves on relatively widely spaced nodes, small calyces, and small, yellow corollas with herkogamous arrangement of stigma and anthers. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 403. | FNA vol. 17, p. 418. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus arenarius, M. floribundus var. subulatus, M. multiflorus, M. subulatus, M. trisulcatus | |
Name authority | (A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) | G. L. Nesom: Phytoneuron 2013-70: 1, figs. 1–5. (2013) |
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