Erythranthe arenaria |
Erythranthe pardalis |
|
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sand-loving monkeyflower |
Pennell's panther |
|
Habit | Annuals, fibrous-rooted or filiform-taprooted. | Annuals, fibrous-rooted or taprooted. |
Stems | erect to ascending, straight or geniculate at nodes, simple or branched, 5–20 cm, villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
decumbent-ascending, erect distally, simple, sometimes branched from proximal to medial nodes, 5–30 cm, short, delicately stipitate-glandular, distals minutely puberulent-glandular, hairs 0.1–0.4 mm (to 1 mm on proximal portions of stems), gland-tipped. |
Leaves | basal and cauline; petiole 0 mm or proximals 1–3(–5) mm; blade 1-veined or palmately 3-veined, elliptic to narrowly elliptic, ovate-elliptic, or ovate-lanceolate, 5–12(–17) × 3–7 mm, base rounded to cuneate-attenuate, margins entire or sparsely dentate to serrate, apex acuminate to acute or obtuse, surfaces villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
usually cauline, basal usually not persistent; petiole: proximals and medials 8–20 mm, distalmost 1–2 mm; blade palmately 3-veined, usually ovate or broadly ovate to depressed-ovate, proximals and medials 7–22 × 6–18 mm, sometimes largest at mid stem, base rounded or cuneate to gradually attenuate, margins shallowly dentate-serrate, teeth 2 or 3(–5) per side mostly distally, apex obtuse to obtuse-acuminate, surfaces sparsely villous to puberulent-glandular, hairs vitreous, gland-tipped, sometimes glabrous. |
Flowers | herkogamous, 1–22, from proximal to distal nodes. |
plesiogamous, 2–12, usually evenly distributed from proximal to distal nodes, chasmogamous, anther pairs in larger corollas slightly separated, stigma at level of distal pair, or both anther pairs and stigma at same level; in smaller corollas without expanded limb and barely exserted beyond calyx margin, both anther pairs and stigma at same level. |
Styles | glabrous. |
glabrous. |
Corollas | yellow, abaxial limb red-dotted, bilaterally symmetric, weakly bilabiate; tube-throat funnelform, 9–12(–14) mm, exserted beyond calyx margin; lobes broadly obovate, apex rounded. |
yellow, throat floor sometimes red-spotted, bilaterally symmetric, bilabiate; tube-throat narrowly funnelform to cylindric, 7–10(–12) mm, exserted 1–3 mm beyond calyx margin; limb expanded 8–12 mm, palate villous. |
Fruiting pedicels | divergent-arcuate, 10–23 mm, villous-glandular, hairs 0.2–0.8 mm, gland-tipped. |
10–35 mm, short, delicately stipitate-glandular, distals minutely puberulent-glandular, hairs 0.1–0.4 mm, gland-tipped. |
Fruiting calyces | usually red-dotted, narrowly campanulate, not or weakly inflated, 5–7(–9) mm, margins distinctly toothed or lobed, villous-glandular, ribs angled, lobes pronounced, erect. |
nodding 45–180º, consistently dark purple-spotted, cylindric-campanulate, inflated, sagittally compressed, 8–11 mm, glabrous or sparsely puberulent-glandular, sometimes minutely hirtellous, throat closing. |
Capsules | included, 4–7 mm. |
included, stipitate, 4–6 mm. |
Anthers | included, glabrous. |
included, glabrous. |
2n | = 32. |
= 28. |
Erythranthe arenaria |
Erythranthe pardalis |
|
Phenology | Flowering May–Sep. | Flowering (Mar–)Apr–May. |
Habitat | Sandy flats, bars, gullies, washes, trails, roadcuts, seasonal creek beds and drainages, rocky slopes, seepy loam, ditches, lake edges, meadows, openings in pine-fir and pine-oak woodlands. | Crevices of serpentine rock, stony red soils, red clay, among boulders, along streams, ditches, tailings at copper mines. |
Elevation | (100–)500–2600(–2800) m. ((300–)1600–8500(–9200) ft.) | 100–700 m. (300–2300 ft.) |
Distribution |
CA
|
CA |
Discussion | Erythranthe arenaria is known from a cluster of six counties of the central Sierra Nevada: Fresno, Madera, Mariposa, Merced, Tulare, and Tuolumne. Most plants of Erythranthe arenaria have relatively even-sized cauline leaves, all sessile to proximally subsessile. Plants in the Yosemite area with persistent basal leaves that are short-petiolate, ovate with a cuneate base, and relatively larger than the more distal cauline ones, and possibly related to E. arenaria, have been named M. floribundus var. subulatus. These might be construed as showing the influence of E. geniculata, but that species occurs only at the lower range of elevation of E. arenaria, while plants referable to Mimulus floribundus var. subulatus occur at least to 2300 m and also have the erect habit characteristic of E. arenaria. These variants should be investigated, especially in the Yosemite area where they appear to be relatively common, with the possibility that they indeed represent a distinct entity. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The relative constancy of Erythranthe pardalis in morphology suggests that genetic influence from other species is slight. It is recognized by its annual duration and relatively delicate habit, ovate to depressed-ovate leaves toothed mostly on the distal margins, small flowers produced from all nodes (proximal to distal), dark-spotted calyces, and stipitate-glandular cauline and foliar vestiture. While the corolla limbs are distinctly expanded, the tubes are only slightly exserted from the calyx, and flowers apparently are plesiogamous. The epithet pardalis alludes to the dark-spotted calyx. Plants of Erythranthe pardalis occur primarily on serpentine rocks and soils but also grow on copper tailings at mine sites. The species is known from Amador, Calaveras, El Dorado, Placer, Tehama, and Tuolumne counties. The plants in Tehama County, geographically and ecologically disjunct from the main range, were recorded as growing in basalt crevices. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 403. | FNA vol. 17, p. 420. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus arenarius, M. floribundus var. subulatus, M. multiflorus, M. subulatus, M. trisulcatus | Mimulus pardalis, M. cupriphilus, M. guttatus var. cupriphilus, M. guttatus var. pardalis |
Name authority | (A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 38. (2012) | (Pennell) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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