Erythranthe alsinoides |
Erythranthe scouleri |
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chickweed monkey-flower, wing-stem monkey-flower |
Columbia River monkey-flower, Scouler's monkey-flower |
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Habit | Annuals, fibrous-rooted. | Perennials, rhizomatous, producing long, sparsely leafy runners from basal nodes. |
Stems | erect, usually simple, (0.5–)2–6(–15) cm, glandular-puberulent, hairs 0.1–0.2 mm, gland-tipped, nodes 2(or 3), usually red-tinged. |
erect, simple or few-branched, 15–80 cm, glabrous. |
Leaves | basal and cauline; petiole 1–20(–30) mm, distinctly 3-veined (winged); blade palmately 3-veined, lanceolate-ovate to ovate, elliptic, or suborbicular, 3–18(–32) × 3–12(–25) mm, base cuneate to truncate, margins dentate to denticulate or subentire, apex acute to obtuse or rounded, surfaces glandular-puberulent, hairs 0.1–0.2 mm, gland-tipped. |
cauline, basal usually not persistent; petiole: proximals and medial 10–25 mm (gradually merging into blade), distals 0 mm; blade palmately (3–)5–7-veined to subpinnate, (proximal to medial) oblong-elliptic to oblong-lanceolate, 25–60 × 8–18 mm, usually 3–4 times longer than wide, base attenuate, margins evenly, shallowly dentate or crenate to mucronate or mucronulate, teeth 10–20 per side, sometimes more deeply toothed at base, apex obtuse to acute, surfaces glabrous. |
Flowers | herkogamous, 1–4(–8), from distal or medial to distal nodes. |
herkogamous, (1 or)2–8, from distal nodes. |
Styles | glabrous. |
minutely, prominently hirsutulous to villosulous. |
Corollas | yellow, abaxial limb with a large maroon splotch, also red-spotted, bilaterally symmetric, ± bilabiate; tube-throat funnelform-cylindric, 6–9 mm, exserted beyond calyx margin; throat open, palate villous, abaxial ridges low. |
yellow, without red markings, bilaterally symmetric, bilabiate; tube-throat funnelform, 20–24 mm, exserted 10–15 mm beyond calyx margin; limb expanded 22–30 mm. |
Fruiting pedicels | 15–32 mm, glandular-puberulent, hairs 0.1–0.2 mm, gland-tipped. |
20–25 mm, glabrous. |
Fruiting calyces | purplish, slightly ridge-angled, campanulate-cylindric, weakly or not inflated, 5–8 mm, margins subtruncate, sparsely minutely stipitate-glandular, lobes 4, (0–)0.5–1 mm, sometimes barely evident, 1 lobe usually slightly longer, margins appearing subtruncate, shallowly convex to rounded-mucronulate. |
ovoid, inflated, sagittally compressed, 13–14 mm, glabrous, throat closing. |
Capsules | included, (3–)5–7 mm. |
unknown. |
Anthers | included, glabrous. |
included, glabrous. |
Erythranthe alsinoides |
Erythranthe scouleri |
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Phenology | Flowering Mar–Jun. | Flowering May–Aug. |
Habitat | Open, rocky slopes, cliff faces, bluffs, mossy rock crevices, ledges, moist rocks, roadsides, along wet paths and trails. | Salt marshes, river banks. |
Elevation | 10–1900 m. (0–6200 ft.) | 0–20 m. (0–100 ft.) |
Distribution |
CA; ID; OR; WA; BC
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OR; WA |
Discussion | Erythranthe alsinoides is distinct in its short, erect stems with few nodes, small, mostly ovate to elliptic-ovate, petiolate leaves, minutely stipitate-glandular vestiture, small corollas with a prominent maroon splotch on the abaxial limb, small, non-inflated mature calyces and, most especially, by its nearly truncate calyx margin. Erythranthe pulsiferae is superficially similar to E. alsinoides but has larger calyces borne on divergent-arcuate pedicels, smaller leaf blades with attenuate to cuneate bases, and the corolla limbs are smaller and lack a prominent maroon splotch. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erythranthe scouleri is distinctive in its oblong-elliptic leaves with long-tapering bases and closely toothed margins, completely glabrous vestiture, long, leafy runners from basal cauline nodes, large corollas with broad limbs, and prominently hairy styles. Several features suggest a close relationship to E. decora, particularly its very large corollas, hairy styles, closely toothed leaf margins, tall, simple, and erect stems, numerous runners, and its geographic range. All collections apparently have been made near the mouth of the Columbia River in Clatsop and Columbia counties, Oregon (G. L. Nesom 2013d). Recent observations (Alexander John Wright, pers. comm.; photos http://www.inaturalist.org/observations/3537008) indicate that it also occurs in Wahkiakum County, Washington, in the Julia Butler Hansen Refuge. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 395. | FNA vol. 17, p. 415. |
Parent taxa | Phrymaceae > Erythranthe | Phrymaceae > Erythranthe |
Sibling taxa | ||
Synonyms | Mimulus alsinoides | Mimulus scouleri, M. guttatus subsp. scouleri |
Name authority | (Douglas ex Bentham) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 37. (2012) | (Hooker) G. L. Nesom: Phytoneuron 2012-39: 44. (2012) |
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