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bushy wallflower, repand wallflower, spreading wallflower, treacle mustard

Pallas' wallflower

Habit Annuals. Perennials; (caudex simple or branched, covered with persistent leaf bases).
Stems

erect, unbranched or branched basally, (0.4–)1.5–4.5(–7) dm.

erect, unbranched, (0.1–)0.4–4dm.

Basal leaves

(often withered by fruiting), similar to cauline.

blade linear to narrowly linear-oblanceolate, 3–5 cm × 1–2 mm, base attenuate, margins entire or denticulate.

Cauline leaves

(proximal and median) petiolate and (distal) sessile, (petiole (0.3–)0.5–2(–3) cm);

blade [linear, narrowly oblanceolate, elliptic, or oblong, (1–)2–8(–11) cm × (2–)5–12(–17) mm, base attenuate], margins sinuate or coarsely dentate to denticulate or repand, (distal) entire or denticulate, (apex acute).

blade reduced distally, often as bracts.

Racemes

considerably elongated in fruit.

(sometimes bracteate basally), elongated in fruit.

Flowers

sepals linear-oblong, 4–6 mm, lateral pair not saccate basally;

petals yellow, narrowly oblanceolate to spatulate, 6–8 × 1.5–2 mm, claw 3–6 mm, apex rounded;

median filaments 4–6 mm;

anthers narrowly oblong, 0.8–1.3 mm.

sepals oblong or narrowly so, 5–9 mm, lateral pair slightly saccate basally;

petals usually purple, rarely lilac, oblanceolate to broadly obovate, 10–20 × 3–5 mm, claw 5–9 mm, apex rounded;

median filaments 6–9 mm;

anthers narrowly oblong, 1–1.5 mm.

Fruiting pedicels

divaricate, stout, as wide as fruit, 2–4(–6) mm.

divaricate to ascending, slender, narrower than fruit, (4–)7–24(–35) mm.

Fruits

widely spreading to divaricate-ascending, narrowly linear, straight or curved upward, somewhat torulose, (2–)3–8(–10) cm × 1.5–2 mm, 4-angled, not striped;

valves with prominent midvein, pubescent outside, trichomes 2-rayed and, fewer, 3-rayed, often glabrous, sometimes pubescent inside;

ovules (40–)50–80(–90) per ovary;

style cylindrical or subclavate, stout, 1–4 mm, sparsely pubescent;

stigma slightly 2-lobed, lobes as long as wide.

erect to divaricate-ascending, narrowly linear, straight or curved inward, not torulose, (3–)5–11(–13) cm × 2–4 mm, latiseptate, not striped;

valves with prominent midvein, pubescent outside, trichomes 2-rayed, glabrous inside;

ovules (28–)36–74 per ovary;

style cylindrical, stout, 0.5–3 mm, sparsely pubescent;

stigma strongly 2-lobed, lobes as long as wide.

Seeds

oblong, 1.1–1.5 × 0.6–0.7 mm;

not winged or, rarely, winged distally.

oblong, 1.7–2.2 × 0.9–0.1.4 mm;

not winged.

Trichomes

of leaves 2-rayed, mixed with fewer 3-rayed ones.

of leaves 2-rayed.

2n

= 16.

= 24, 36.

Erysimum repandum

Erysimum pallasii

Phenology Flowering Apr–Jun. Flowering Jun–Aug.
Habitat Disturbed sites, roadsides, fields, waste places, barren hillsides, brush communities, pastures Cliffs, shaley slopes, talus, sandy ground, alpine areas, clay banks, pebbles, gravel
Elevation 0-2100 m (0-6900 ft) 0-1300 m (0-4300 ft)
Distribution
from FNA
AL; AR; AZ; CA; CO; IA; ID; IL; IN; KS; KY; LA; MA; MD; MI; MO; MS; MT; NC; NE; NJ; NM; NV; NY; OK; OR; PA; TN; TX; UT; VA; WA; WV; WY; BC; ON; QC; Eurasia; n Africa [Introduced in North America; introduced also in South America, Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; NT; NU; YT; Greenland; e Asia (Russian Far East)
[BONAP county map]
Discussion

R. C. Rollins (1993) and G. A. Mulligan (2002) listed haploid chromosome numbers of n = 12, 14, 18, and 21 for Erysimum pallasii, but it is doubtful that those deviating from x = 6 are correct. Tetraploid counts (2n = 24) are known from both Canada and Russian Far East, whereas hexaploid counts are known only from the latter region. G. B. Rossbach (1958) and Rollins recognized plants with racemes bracteate to the middle as var. bracteosum, but this distinction is rather artificial, and the species shows considerable variation in the number of bracteate flowers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 544. FNA vol. 7, p. 543.
Parent taxa Brassicaceae > tribe Erysimeae > Erysimum Brassicaceae > tribe Erysimeae > Erysimum
Sibling taxa
E. ammophilum, E. arenicola, E. asperum, E. capitatum, E. cheiranthoides, E. cheiri, E. coarctatum, E. concinnum, E. franciscanum, E. hieraciifolium, E. inconspicuum, E. insulare, E. menziesii, E. occidentale, E. pallasii, E. perenne, E. suffrutescens, E. teretifolium
E. ammophilum, E. arenicola, E. asperum, E. capitatum, E. cheiranthoides, E. cheiri, E. coarctatum, E. concinnum, E. franciscanum, E. hieraciifolium, E. inconspicuum, E. insulare, E. menziesii, E. occidentale, E. perenne, E. repandum, E. suffrutescens, E. teretifolium
Synonyms Cheirinia repanda Cheiranthus pallasii, Cheiranthus pygmaeus, E. pallasii var. bracteosum, E. pygmaeum, Hesperis hookeri
Name authority Linnaeus: Demonstr. Pl., 17. (1753) (Pursh) Fernald: Rhodora 27: 171. (1925)
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