Erysimum perenne |
Erysimum suffrutescens |
|
---|---|---|
sand dune wallflower, Sierra wallflower |
island wallflower, suffrutescent wallflower |
|
Habit | Perennials or, rarely, biennials; (caudex slender). | Perennials or subshrubs. |
Stems | erect, unbranched or branched (few to several) basally, 0.4–6.5 dm. |
ascending, proximal branches terminating in sterile rosettes, (woody at base), 1.5–8.1 dm. |
Basal leaves | blade spatulate to broadly oblanceolate, 2.5–7 cm × 3–10 mm, base attenuate, margins dentate or subentire, apex often obtuse, (surfaces pubescent adaxially, trichomes 2 or 3–5-rayed). |
blade linear to linear-oblanceolate, 3.7–15 cm × 1.5–6(–7) mm, base attenuate, margins usually entire, rarely sparsely denticulate, apex acute. |
Cauline leaves | (distal) sessile; blade margins often entire. |
(distal) sessile; blade similar to basal. |
Racemes | considerably elongated in fruit. |
considerably elongated in fruit. |
Flowers | sepals linear-oblong to oblong, 8–12 mm, lateral pair saccate basally; petals yellow, broadly obovate to suborbicular, 15–22 × 3.5–6 mm, claw 8–14 mm, apex rounded; median filaments 7–14 mm; anthers linear, 3–4 mm. |
sepals oblong to linear-oblong, 6–11 mm, lateral pair saccate basally; petals yellow, obovate to suborbicular, (11–)14–20(–22) × (3–)4–11.5 mm, claw 8–13 mm, apex rounded; median filaments 7–11 mm; anthers linear, 2.5–4 mm. |
Fruiting pedicels | divaricate-ascending, slender, narrower than fruit, 4–12 mm. |
ascending, slender, narrower than fruit, (3–)5–10 mm. |
Fruits | erect to ascending, narrowly linear, straight, torulose, 3.8–14 cm × 1.2–3 mm, latiseptate, not striped; valves with prominent midvein, pubescent outside, trichomes 2 or 3 (or 4)-rayed, glabrous inside; ovules 26–44 per ovary; style cylindrical, slender, (1.5–)2–5.5 mm, sparsely pubescent; stigma subentire to slightly 2-lobed, lobes as long as wide. |
ascending to spreading, narrowly linear, straight or only slightly curved inward, not torulose, (2–)3–8.4(–11) cm × 1.5–2.4(–3.5) mm, 4-angled to only slightly latiseptate, not striped; valves with prominent midvein, pubescent outside, trichomes 2–4-rayed, glabrous inside; ovules 48–82 per ovary; style cylindrical, stout, 0.5–4 mm, sparsely pubescent; stigma slightly 2-lobed, lobes as long as wide. |
Seeds | ovoid, 2–3.4 × 1–2 mm; not winged or, rarely, winged distally. |
oblong, 1.5–2.5(–3) × (0.7–)1–1.2(–1.4) mm; not winged or winged apically. |
Trichomes | of leaves 2–5-rayed. |
of leaves 2-rayed, sometimes mixed with 3- or 4-rayed ones. |
2n | = 36. |
= 36. |
Erysimum perenne |
Erysimum suffrutescens |
|
Phenology | Flowering Jun–Sep. | Flowering Dec–Aug. |
Habitat | Alpine fellfields, decomposing marble, gravelly ground and knolls, rocky slopes, talus, granitic sand | Stabilized coastal sand dunes, coastal scrub vegetation |
Elevation | 2000-4000 m (6600-13100 ft) | 0-150 m (0-500 ft) |
Distribution |
CA; NV; OR
|
CA
|
Discussion | Erysimum perenne is a high alpine species of the western sierras in California from Fresno, Inyo, and Madera counties northward into Plumas, Siskiyou, and Trinity counties. Its range in Nevada appears to be restricted to Douglas and Washoe counties. The limits of Erysimum perenne have been controversial, and it is with some hesitation that I recognize it as a species. G. B. Rossbach (1958) accepted it as a distinct species, R. A. Price (1993) transferred it (invalidly) to a subspecies of E. capitatum, R. C. Rollins (1993) treated it as a variety of E. capitatum, and N. H. Holmgren (2005b) treated the name as a synonym of E. capitatum. It is readily distinguished from E. capitatum by having torulose (versus not torulose) and flattened (versus 4-angled or flattened) fruits, slender (versus stout or, rarely, slender) and longer styles (1.5–)2–5.5 mm (versus 0.2–2.5(–3) mm), and yellow (versus orange to, rarely, yellow) petals. Where the two species are allopatric, they remain consistently distinct, but at lower elevations, where their ranges overlap, the distinction becomes blurred. In such areas of overlap, one finds fruit variation ranging from distinctly torulose to non-torulose, as well as continuity in the other characters above. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Erysimum suffrutescens is restricted to the coastal regions of Los Angeles County northward into San Luis Obispo, Santa Barbara, and Ventura counties. Both G. B. Rossbach (1958, 1958b) and R. C. Rollins (1993) treated it as a distinct species, but R. A. Price (1993) transferred it (invalidly) to a subspecies of E. insulare. The latter species has angustiseptate (versus 4-angled to slightly latiseptate) fruits, and the two are sufficiently distinct to be recognized as independent species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 543. | FNA vol. 7, p. 544. |
Parent taxa | Brassicaceae > tribe Erysimeae > Erysimum | Brassicaceae > tribe Erysimeae > Erysimum |
Sibling taxa | ||
Synonyms | E. asperum var. perenne, Cheiranthus perennis, Cheirinia nevadensis, E. capitatum var. perenne, E. nevadense | Cheiranthus suffrutescens, E. concinnum subsp. suffrutescens, E. suffrutescens var. grandifolium |
Name authority | (S. Watson ex Coville) Abrams: in L. Abrams and R. S. Ferris, Ill. Fl. Pacific States 2: 318. (1944) | (Abrams) Rossbach: Aliso 4: 121. (1958) |
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