Erysimum concinnum |
Erysimum |
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coast wallflower, curly wallflower, headland wallflower, Pacific wallflower |
wallflower |
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Habit | Biennials or perennials; (short-lived). | Plants not scapose; pubescent, trichomes sessile, medifixed, appressed, 2-rayed (malpighiaceous) or 3–5(–8)-rayed (stellate), rays (when 2) parallel to long axis of stems, leaves, sepals, and fruits. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | erect, unbranched or branched distally, 0.4–5(–7) dm. |
erect or ascending [decumbent], unbranched or branched basally and/or distally. |
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Leaves | basal and cauline; petiolate or sessile; basal rosulate or not, petiolate, blade margins usually entire, dentate, sinuate-dentate, or denticulate, rarely pinnatifid or pinnatisect; cauline petiolate or sessile, blade (base cuneate or attenuate [auriculate]), margins entire, dentate, denticulate, dentate-sinuate, or repand. |
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Basal leaves | blade (slightly fleshy), spatulate to oblanceolate, 2–11 cm × 4–20 mm, base attenuate, margins sinuate-dentate to coarsely dentate, apex rounded to subacute. |
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Cauline leaves | (distal) sessile; blade margins entire or denticulate. |
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Trichomes | of leaves 2- or 3(–7)-rayed. |
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Racemes | considerably elongated in fruit. |
(densely flowered, E. pallasii bracteate basally). |
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Flowers | sepals oblong, 8–19 mm, lateral pair saccate basally; petals yellow to cream, suborbicular to broadly obovate, 15–32 × 6–16 mm, claw 8–12 mm, apex rounded; median filaments 8–11 mm; anthers linear, 3–4 mm. |
sepals oblong or linear, lateral pair saccate or not basally (pubescent); petals suborbicular, obovate, or spatulate, claw differentiated from blade (subequaling or longer than sepals, apex rounded [emarginate]); stamens (erect), tetradynamous; filaments not dilated basally; anthers oblong or linear; nectar glands (1, 2, or 4), distinct or confluent, subtending bases of stamens, median glands present or absent. |
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Fruiting pedicels | ascending, stout, narrower than fruit, 2–4(–6) mm. |
erect, ascending, divaricate, reflexed, horizontal, or spreading, slender or stout (nearly as wide as fruit). |
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Fruits | usually ascending to suberect, rarely divaricate-ascending, narrowly linear, straight or curved inwards, not torulose, (3–)5–13 cm × 2.2–5 mm, terete when immature, becoming strongly latiseptate, not striped; valves with obscure midvein, pubescent outside, trichomes 2–5-rayed, glabrous inside; ovules 42–68 per ovary; style cylindrical or flattened, stout, 0.5–2.5 mm, sparsely pubescent; stigma 2-lobed, lobes as long as wide. |
usually sessile, rarely shortly stipitate (gynophore to 4 mm), usually linear or narrowly so [oblong], smooth or torulose, (keeled or not); valves each with obscure to prominent midvein, pubescent outside, usually glabrous inside; replum rounded; septum complete, (not veined); ovules [15–]20–120 per ovary; (style relatively short, rarely 1/2 as long as or subequaling fruit, often pubescent); stigma capitate. |
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Seeds | broadly ovate to suborbicular, (1.5–)2–4 × 1.5–3 mm; wing continuous. |
plump or flattened, winged, margined, or not winged, oblong, ovoid, obovate, or suborbicular; seed coat (minutely reticulate), mucilaginous when wetted; cotyledons incumbent, rarely accumbent. |
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x | = (6) 7, 8 (9–17). |
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2n | = 36. |
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Erysimum concinnum |
Erysimum |
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Phenology | Flowering Mar–Jun. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Coastal bluffs, dunes, prairies | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 0-400 m (0-1300 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
CA; OR
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North America; n Mexico; Central America; Europe; Asia; n Africa; Atlantic Islands (Macaronesia) [Introduced in South America, Australia] |
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Discussion | Erysimum concinnum is a coastal species known from Curry County in Oregon, and from Del Norte, Humboldt, Marin, Mendocino, and Sonoma counties in California. Both G. B. Rossbach (1958) and R. C. Rollins (1993) treated it as a distinct species, but R. A. Price (1993) reduced it (invalidly) to a subspecies of E. menziesii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 150 (19 in the flora). Erysimum is found in the northern hemisphere, primarily Asia and Europe, with eight species in northern Africa and Macaronesia, and one each endemic to Baja California (E. moranii Rollins) and Costa Rica and Guatemala (E. ghiesbreghtii J. D. Smith). Of the 21 species found in North America, four are naturalized. Most of the native species have x = 9 and are believed to represent a monophyletic group (R. A. Price 1987). Erysimum is a taxonomically difficult genus much in need of comprehensive phylogenetic and systematic studies covering its entire range. The principal sources of difficulty are the inflation in the number of species described, the heavy reliance on vegetative morphological characters in the delimitation of species, and the inadequacy of most herbarium specimens. In order to reliably identify a given sample, one often needs a complete specimen that has basal leaves, flowers, mature fruits, and seeds. Unfortunately, plants of most species shed their basal leaves or have no flowers when at full fruit maturity. Another complicating factor in North America is that almost all of the native species readily hybridize in areas of overlap to produce wide arrays of intermediates that backcross with the parents and blur species boundaries. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 540. | FNA vol. 7, p. 534. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Cheiranthus, Cheirinia, Cuspidaria, Syrenia | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | Eastwood: Zoë 5: 103. (1901) | Linnaeus: Sp. Pl. 2: 660. (1753): Gen. Pl. ed. 5, 296. (1754) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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