bastard-sage, bastard-sage buckwheat, Wright's buckwheat, Wright's wild buckwheat
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California buckwheat, California wild buckwheat, eastern Mojave buckwheat, eastern Mojave wild buckwheat
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Shrubs, subshrubs, or herbs, rarely scapose, (1–)1.5–10 × 1–15(–18) dm or, if matted, 0.1–2.5(–3) × 0.5–3(–5) dm, , lanate to thinly tomentose, or glabrous, grayish to greenish or reddish. |
Shrubs or subshrubs, compact to spreading or rounded and more or less erect, occasionally decumbent, infrequently scapose, (1–)2–15 × 2–25(–30) dm, tomentose to canescent, floccose, or glabrous. |
spreading to erect, with or without persistent leaf bases, up to 1/2 or more height of plant; caudex stems absent or spreading, occasionally matted; aerial flowering stems erect to spreading, stout to slender, solid, not fistulose, (0.1–)0.5–4(–6) dm, tomentose, floccose, or glabrous. |
sprawling or spreading to erect, often with persistent leaf bases, up to 1/2 or more height of plant; caudex stems absent or matted to spreading; aerial flowering stems erect to spreading, slender, solid, not fistulose, 0.3–2.5(–3) dm, tomentose, canescent, or glabrous. |
basal and fasciculate in terminal tufts, or cauline and fasciculate, occasionally 1 per node; petiole 0.02–0.5(–1) cm, tomentose to floccose; blade oblanceolate to broadly elliptic, 0.1–3 × 0.1–1 cm, tomentose to floccose, sometimes subglabrous or glabrous and green adaxially, margins plane, sometimes revolute. |
cauline, 1 per node or fasciculate; petiole 0.1–0.3 cm, canescent; blade linear to linear-oblanceolate or oblanceolate, 0.6–1.5(–1.8) × 0.05–0.4(–0.6) cm, white-tomentose or canescent to subglabrous abaxially, tomentose or canescent and grayish, subglabrous, or glabrous and green adaxially, margins often revolute. |
virgate or cymose with involucres disposed at tips racemosely arranged involucres, rarely capitate, (1–)5–20 × (1–)10–40 cm; branches dichotomous, tomentose, floccose, or glabrous; bracts 3, triangular, scalelike, 0.5–3.5 mm. |
cymose, infrequently cymose-umbellate or capitate, compact to open, occasionally flat-topped, 0.2–20 × 0.2–15 cm; branches dichotomous, infrequently absent, tomentose to canescent or glabrous; bracts usually 3, scalelike, triangular, and 1–3 mm, or leaflike, linear to oblanceolate, and 3–10 × 1–3 mm. |
absent. |
absent. |
1 per node, turbinate to narrowly campanulate, (0.7–)1–4 × 1–2.5 mm, tomentose, floccose, or glabrous; teeth 5, erect, 0.3–1 mm. |
(1–)3–8 per cluster, turbinate to campanulate, 2–4 × 1.5–3 mm, canescent, pubescent, glabrous, or subglabrous; teeth 5, erect, 0.3–1.2 mm. |
1–4 mm; perianth white to pink or rose, glabrous; tepals connate proximal 1/4, monomorphic, obovate; stamens exserted, 1.5–4 mm; filaments glabrous or sparsely pilose proximally. |
2.5–3 mm; perianth white to pinkish, glabrous or pubescent; tepals connate proximal 1/4, monomorphic, usually elliptic to obovate; stamens exserted, 2.5–5 mm; filaments subglabrous or pubescent proximally. |
light brown to brown, (1–)1.5–3 mm, glabrous. |
light brown to brown, 1.8–2.5 mm, glabrous. |
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AZ; CA; NM; NV; TX; UT; including nw Mexico
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AZ; CA; NV; UT; including nw Mexico
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Varieties 9 (6 in the flora). Eriogonum wrightii is subdivided into several varieties most of which are distinct, although a few have rather indistinct boundaries. Some of the variation has yet to be fully resolved, especially in the var. nodosum complex where, at least in Mexico, one additional expression remains to be named. A clear distinction between var. subscaposum and E. kennedyi is not possible in southern California (see discussion below). Nearly all of the varieties of E. wrightii are in cultivation, although the most elegant (var. olanchense) has yet to be so honored. Few ethnobotanical uses of bastard-sage are reported in the literature. L. C. Wyman and S. K. Harris (1951) noted that the Kayenta Navajo use it (var. wrightii) as an emetic, while M. L. Zigmond (1981) stated that the Kawaiisu used the pounded seeds (probably of var. subscaposum) in a beverage or as a dry meal. Members of the species are food plants for the rare Rita dotted-blue butterfly (Euphilotes rita), the Pacific dotted-blue (E. enoptes), the veined blue (Plebeius neurona), and the Mormon metalmark (Apodemia mormo mormo). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 5 (4 in the flora). Eriogonum fasciculatum is a complex, polyploid series of variants that are generally distinct but often difficult to distinguish morphologically. Variety emphereium Reveal is confined to central Baja California, Mexico. The introduction of Eriogonum fasciculatum as a decorative roadside plant by the California Department of Transportation is resulting in hybrid populations involving E. cinereum. The aggressively weedy and (for Arizona) exotic variety foliosum is rapidly invading the native habitat of var. polifolium. Members of E. fasciculatum are food plants for several butterflies, notably the Bernardino dotted-blue (Euphilotes bernardino), lupine blue (Plebeius lupini), Mormon metalmark (Apodemia mormo), and Behr’s metalmark (A. virgulti). Probably the butterfly most commonly seen with the species is the nut-brown hairstreak (Satyrium saepium), which frequents plants in full flower. Eriogonum fasciculatum is also the most important native source of honey in California. This widespread species was used extensively by Native Americans for a variety of ailments. Its application for pain and headaches (D. P. Barrows 1900; K. Hedges 1986; E. W. Voegelin 1938) was rather common, as was its general use for diarrhea (Hedges; Voegelin). M. L. Zigmond (1981) reported that the Kawaiisu lined their acorn granaries with leaves of var. proliferum to keep out rain—a daunting challenge given the size of the leaves! L. Hinton (1975) reported the use of a decoction of dried flowers and roots to maintain a healthy heart, and M. C. Stevenson (1915) indicated that a powder derived from the roots was used by the Zuñi to treat wounds, whereas a root decoction was taken for colds and hoarseness. B. R. Bocek (1984) reported that the Costanoan Indians of California used a decoction of the plant to treat unspecified urinary problems. F. H. Elmore (1943) reported the use of a decoction of var. proliferum by the Navajo (Diné) people as an anti-witchcraft medicine. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Plants loosely to compactly matted or herbs | → 2 |
1. Plants shrubs or subshrubs | → 3 |
2. Plants 0.5-2.5(-3) dm; leaf blades 0.5-1(-1.2) cm; involucres 1.5-4 mm; e and s California,wc Nevada | var. subscaposum |
2. Plants 0.1-0.3(-0.6) dm; leaf blades 0.1-0.25 cm; involucres 0.8-1.7(-2) mm; se California | var. olanchense |
3. Flowering stems and branches grayish, lanate to densely tomentose; sw Arizona, se California | var. nodosum |
3. Flowering stems and inflorescence branches whitish, reddish, or greenish, tomentose to floccose; widespread | → 4 |
4. Petiole bases forming distinct ring around stem; leaf blades 0.2-0.6(-1) × 0.1-0.3(-0.4) cm; s California | var. membranaceum |
4. Petiole bases not forming distinct ring around stem; leaf blades 0.5-3 × 0.2-1 cm; sw United States, Mexico | → 5 |
5. Leaf blades 0.5-1.5 × 0.2-0.5(-0.7) cm; involucres 2-2.5 mm; flowers 2.5-3.5 mm; se California to w Texas | var. wrightii |
5. Leaf blades 1.5-3 × 0.5-1 cm; involucres 3-4 mm; flowers 3-4 mm; c and nw California | var. trachygonum |
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1. Leaves thinly white-tomentose abaxially, glabrous adaxially; perianths and involucres mostly glabrous; plants usually decumbent; primarily coastal 80c. Eriogonum fasciculatum var. fasciculatum | → 1 |
1. Leaves, perianths, and involucres pubescent or, if subglabrous, then plants of desert regions; plants erect to rounded; generally inland | → 2 |
2. Leaves light yellow-green, subglabrous adaxially; involucres and perianths glabrous or subglabrous | var. flavoviride |
2. Leaves dark green or grayish, usually pubescent adaxially; involucres and perianths pubescent | → 3 |
3. Inflorescences capitate to cymose-umbellate, rarely cymose; leaves canescent on both surfaces or densely tomentose abaxially and canescent adaxially; leaf margins plane or infrequently revolute; mainly deserts | var. polifolium |
3. Inflorescences open and mostly cymose; leaves densely white-tomentose abaxially, less so to green and floccose adaxially; leaf margins usually tightly revolute; mainly inner coastal foothills | var. foliolosum |
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FNA vol. 5, p. 289. |
FNA vol. 5, p. 297. |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. zionis |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
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E. trachygonum subsp. wrightii |
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Torrey ex Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 14: 15. (1856) |
Bentham: Trans. Linn. Soc. London 17: 411. (1836) |
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