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Wooton's buckwheat, Wooton's wild buckwheat

marum-leaf buckwheat, marum-leaf wild buckwheat, mountain buckwheat

Habit Herbs, erect, 4–5 × 3–5 dm, tomentose to floccose. Herbs, matted, dioecious, (0.1–)0.5–5 × 2–8 dm, floccose or glabrous.
Stems

caudex absent;

aerial flowering stems erect, stout, solid, not fistulose, arising directly from a taproot, 1–2.5 dm, tomentose to floccose.

caudex spreading;

aerial flowering stems erect, slender, solid, not fistulose, arising at nodes of caudex branches and at distal nodes of short, non-flowering aerial branches, (0.1–)0.5–4 dm, floccose or glabrous.

Leaves

basal, not in rosettes;

petiole 2–5 cm, tomentose to floccose;

blade elliptic, (2.5–)3–6.5 × 1.5–3 cm, densely whitish- to tannish-tomentose abaxially, thinly tomentose to floccose and greenish adaxially, margins entire, plane.

in loose to compact basal rosettes;

petiole 0.3–2 cm, often tomentose;

blade ovate to oval, 0.3–3 × 0.3–1 cm, densely tannish- to brownish-lanate abaxially, glabrate and green to olive green adaxially, margins entire, plane.

Inflorescences

compound-umbellate, 5–15 × 5–20 cm;

branches tomentose to floccose;

bracts 3–6, leaflike at proximal node, 2–4(–5) × 0.7–1.8 cm, often scalelike distally.

capitate, 0.5–2 cm wide, mature pistillate plants open and elongate-umbellate, 1–5 × 1–7 cm;

branches usually glabrous;

bracts 5–10, leaflike, 0.2–0.8 cm, often absent immediately below involucre.

Involucres

1 per node, campanulate, 3–6 × 3–7 mm, thinly floccose;

teeth 5–7, erect, 0.3–0.6 mm.

1 per node but occasionally appearing congested, turbinate and 2–3 × 1.5–2 mm or campanulate and 4–5 × 5–7 mm, sparsely tomentose or glabrous;

teeth 5–6, erect, 0.4–0.7 mm.

Flowers

4–7(–9) mm, including 1–1.5 mm stipelike base;

perianth bright yellow, densely pubescent abaxially;

tepals dimorphic, those of outer whorl lanceolate to elliptic, 3–6 × 1.5–2.5, those of inner whorl elliptic, 5–9 × 2–3.5 mm;

stamens exserted, 3–5 mm;

filaments pilose proximally.

stipelike base 0.5–1 mm;

perianth yellow, glabrous; staminate flowers 1.5–3 mm, tepals ovate; pistillate flowers 4–7 mm, tepals oblanceolate, often becoming reddish in fruit;

stamens exserted, 2–3 mm;

filaments pilose proximally.

Achenes

light brown to brown, 4–8 mm, glabrous except for sparsely pubescent beak.

light brown to brown, 3.5–5 mm, glabrous except for sparsely pubescent beak.

2n

= 32.

Eriogonum wootonii

Eriogonum marifolium

Phenology Flowering Jul–Sep.
Habitat Sandy to gravelly slopes, sagebrush communities, montane conifer woodlands
Elevation 2000-2800 m (6600-9200 ft)
Distribution
from FNA
NM
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV; OR; WA
[WildflowerSearch map]
[BONAP county map]
Discussion

Eriogonum wootonii is known only from the San Francisco Mountains of Lincoln and Otero counties in southern New Mexico. It is related to, but significantly disjunct from, E. arcuatum, and is part of a three-species complex that includes E. correllii of northern Texas and E. allenii of Virginia and West Virginia, and that skips across the central United States, the result of fragmentation associated with the southward push of Pleistocene glaciation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Eriogonum marifolium is dioecious, with the two sexes having such markedly different morphology that collectors often consider each to represent a different species. In Oregon, male plants of E. marifolium and E. umbellatum var. haussknechtii can be difficult to distinguish. In El Dorado and Placer counties, California, herbarium specimens of E. marifolium and E. incanum can be difficult to separate, as their distribution ranges overlap there.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Involucres turbinate, 2-3 × 1.5-2 mm; ne California, wc Nevada, s Oregon, disjunct to Yakima County, Washington
var. marifolium
1. Involucres campanulate, 4-5 × 5-7 mm; nc California
var. cupulatum
Source FNA vol. 5, p. 370. FNA vol. 5, p. 365.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Oligogonum Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Oligogonum
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. marifolium var. cupulatum, E. marifolium var. marifolium
Synonyms E. jamesii var. wootonii E. cupulatum, E. marifolium var. apertum
Name authority (Reveal) Reveal: Harvard Pap. Bot. 9: 206. (2003) Torrey & A. Gray: Proc. Amer. Acad. Arts 8: 161. (1870)
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