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bright green buckwheat, two-tooth wild buckwheat, twotooth buckwheat

Twisselmann's buckwheat, Twisselmann's wild buckwheat

Habit Herbs, spreading, annual, (0.5–)1–2(–3) dm, tomentose, greenish, grayish, or reddish. Herbs, matted, occasionally polygamodioecious, (0.2–)0.5–1.5 × 3–4 dm, tomentose.
Stems

caudex absent;

aerial flowering stems erect, not striated or angled, solid, not fistulose, 0.2–0.8 dm, tomentose.

caudex spreading;

aerial flowering stems erect or nearly so, slender, solid, not fistulose, arising at nodes of caudex branches and at distal nodes of short, nonflowering aerial branches, (0.2–)0.5–1.2 dm, tomentose, with a whorl of 4–8 leaflike bracts ca. midlength, similar to leaf blades, 0.4–0.8 × 0.1–0.3 cm.

Leaves

basal and cauline;

basal: petiole 0.5–1 cm, floccose, blade lanceolate to obovate, (0.5–)2–3 × 1.5–2 cm, tomentose abaxially, floccose to glabrate and grayish to greenish adaxially, margins entire or occasionally crenulate;

cauline sessile, blade lanceolate to oblanceolate, 0.5–2 × 0.3–1 cm, similar to basal blade.

in loose to congested basal rosettes;

petiole 0.2–0.4 cm, thinly tomentose;

blade oblanceolate to elliptic, 0.5–1 × (0.2–)0.4–0.6 cm, white- or grayish-tomentose abaxially, thinly tomentose to nearly glabrous and dull green adaxially.

Inflorescences

cymose, open, 4–25 × 5–30 cm;

branches tomentose;

bracts 3, scalelike, 0.5–2.5 × 1–2 mm.

capitate, 0.7–1.5 cm wide;

branches absent;

bracts absent immediately below involucre.

Peduncles

spreading, straight, capillary, 1–2 cm, sparsely glandular.

Involucres

campanulate, 2–3 × 2–4 mm, finely glandular;

teeth 5, erect, 0.4–0.8 mm.

1 per node, campanulate, (2.5–)4–5 × 4–6 mm;

teeth 6–9, lobelike, strongly reflexed, (2–)3–5 mm.

Flowers

1–2.5 mm;

perianth white to rose, glandular;

tepals dimorphic, those of outer whorl obovate to spatulate, broadly inflated above middle, those of inner whorl narrowly spatulate;

stamens included, 1–2 mm;

filaments pilose proximally.

(4–)5–6 mm, including 1–1.3 mm stipelike base;

perianth pale yellow, villous abaxially;

tepals slightly dimorphic, those of outer whorl obtuse, 3–5 × 3–3.5 mm, those of inner whorl broadly oblanceolate, 5–6 × 2.5–3 mm;

stamens exserted, 4–5 mm;

filaments pilose proximally.

Achenes

light brown, 3-gonous, 1–1.5 mm, glabrous.

light brown, 5–5.5 mm, glabrous except for sparsely pubescent beak.

Eriogonum viridescens

Eriogonum twisselmannii

Phenology Flowering Apr–Nov. Flowering Jun–Sep.
Habitat Sandy to gravelly or clayey flats and slopes, mixed grassland, chaparral, saltbush, and creosote bush communities, oak and conifer woodlands Rocky outcrops, montane conifer woodlands
Elevation 100-1700 m (300-5600 ft) 2300-2600 m (7500-8500 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Eriogonum viridescens is infrequent in the Inner Coast and the Transverse ranges (Fresno, Kern, Kings, Los Angeles, Merced, Monterey, Riverside, San Benito, western San Bernardino, San Luis Obispo, Santa Barbara, and Ventura counties). Unlike those of the more widespread E. maculatum, the outer tepals of E. viridescens are swollen apically, rather than basally. A collection made at Crystal Springs in the Coso Mountains of Inyo County (Coville & Funston 909, DS, US) probably is accurately labeled but the location remains to be confirmed by modern collections.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Eriogonum twisselmannii is a rare species of the southern Sierra Nevada and is known only from Slate Mountain and The Needles in Tulare County. It is worthy of being on a “watch” list of rare plants in California, but because of its remote location, it is not threatened.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 412. FNA vol. 5, p. 364.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Oligogonum
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Synonyms E. angulosum subsp. bidentatum, E. angulosum subsp. viridescens, E. angulosum var. viridescens, E. bidentatum E. douglasii var. twisselmannii
Name authority A. Heller: Muhlenbergia 2: 25. (1905) (J. T. Howell) Reveal: in P. A. Munz, Suppl. Calif. Fl. 40. (1968)
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