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dogtongue buckwheat, sandhill wild buckwheat

fine branch wild buckwheat, sand buckwheat, sand wild buckwheat

Habit Herbs, 5–12 × 2–5 dm, white- to rufous-tomentose. Shrubs, erect to spreading, not scapose, (2–)3–10(–12) × 5–15(–20) dm, white-tomentose to floccose, or green and nearly glabrous.
Stems

spreading, often without persistent leaf bases, up to 1/2 height of plant;

caudex stems absent or spreading in moving sand;

aerial flowering stems spreading, slender, solid, not fistulose, 0.3–1 dm, white-tomentose, floccose to glabrate or glabrous.

Aerial flowering stems

erect, 1.5–7 dm, rounded and smooth.

Leaves

petiole 2–7 cm;

blade oblong to elliptic or spatulate, 4–15 × 1.5–4 cm, densely tomentose abaxially, subglabrous or glabrous adaxially.

cauline, 1 per node;

petiole 0.2–0.5 cm, tomentose to floccose;

blade linear-lanceolate to linear-oblanceolate to narrowly oblong, or lanceolate to narrowly elliptic, 1.5–4 × 0.2–0.8(–1.2) cm, densely white-tomentose abaxially, less so and greenish adaxially, margins plane, infrequently revolute.

Inflorescences

(3–)5–30 × 3–20 cm;

bracts 3–5, leaflike, ovate to elliptic, 1–6 × 0.5–2.5 cm proximally, semileaflike, usually elliptic, 4–30 × 1–20 mm distally.

cymose, open, 5–40 × 10–50 cm;

branches dichtomous proximally, often with involucres racemosely arranged proximally, tomentose to floccose or glabrate, occasionally glabrous;

bracts 3, scalelike, linear to triangular, 1–3(–6) mm.

Peduncles

absent.

absent.

Involucres

turbinate-campanulate to campanulate, 3–4.5(–5) × 3–5 mm;

teeth 0.1–0.5 mm.

1 per node, turbinate to turbinate-campanulate, 1.5–3 × 1–2 mm, tomentose to floccose or glabrous;

teeth 5, erect, 0.4–0.7 mm.

Flowers

8–13 mm, including 2.5–3.5(–4) mm stipelike base;

perianth cream to light tan, densely tannish- to rusty-lanate abaxially;

tepals dimorphic, those of outer whorl broadly lanceolate, 3–5 × 1.7–2.2 mm, those of inner whorl obovate to nearly orbiculate, 3–6 × 3–4 mm;

stamens 3.5–5 mm;

filaments pilose.

(2–)2.5–3.5 mm;

perianth white or pale yellow to yellow, glabrous;

tepals connate proximal 1/4–1/3, essentially monomorphic, oblong to broadly obovate;

stamens slightly exserted, 2–4 mm;

filaments sparsely pilose proximally.

Achenes

brown, 4.5–5 mm, glabrous.

light brown, 2.5–3.5 mm, glabrous except for minutely papillate beak.

Eriogonum tomentosum

Eriogonum leptocladon

Phenology Flowering May–Oct.
Habitat Sandy soil, mixed grassland communities, pine and oak woodlands
Elevation 0-100(-200) m (0-300(-700) ft)
Distribution
from FNA
AL; FL; GA; SC
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; NM; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Eriogonum tomentosum is widespread in a series of somewhat disjunct populations mainly on the coastal plain. It supposedly is found in North Carolina, but no specimens have been seen. An unnumbered J. K. Small collection from Miami, Florida (Nov. 1904, NY), surely is mislabeled. A Cuthbert collection (30 Jul. 1876, MICH) from Augusta may be from a garden, as that location is well out of the known range of the species. Mark Catesby made the first collections (BM-Sloane, OXF) of the species in the 1720s. T. Walter (1788) ascribed his specimen (BM-Walter) to a European species of Saxifragaceae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Eriogonum leptocladon is localized on moving sands on the Colorado Plateau. It is not unusual, therefore, for portions of the shrub to be buried in sand, especially the main, woody trunk from which the flowering stems arise. In the Four Corners area three species share portions of overlapping ranges, and herbarium material from there can be difficult to separate. In northwestern New Mexico, var. ramosissimum approaches E. effusum, and in eastern San Juan County the two can be distinguished only by the open, spreading inflorescence branching pattern of var. ramosissimum as compared to the densely branched, compact inflorescence of E. effusum. In the field, the former is confined to sandy areas whereas the latter is typically on heavier, usually clayey soils. In south-central Utah, var. ramosissimum approaches E. nummulare. There, both are found on moving sands, and only the degree of stoutness of the branches can be used to differentiate them when cauline leaves are absent. For the most part, the inflorescence branches of E. nummulare are more rigid and stouter than those of var. ramosissimum, which tend to be flexible and slender. When leaves are present, those of the former are broader and more apically rounded compared to those of the latter.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Perianths pale yellow to yellow
var. leptocladon
1. Perianths white
→ 2
2. Flowering stems and inflorescence branches tomentose, rarely floccose
var. ramosissimum
2. Flowering stems and inflorescence branches glabrous
var. papiliunculi
Source FNA vol. 5, p. 330. FNA vol. 5, p. 249.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eriogonum Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. leptocladon var. leptocladon, E. leptocladon var. papiliunculi, E. leptocladon var. ramosissimum
Synonyms E. effusum subsp. leptocladon
Name authority Michaux: Fl. Bor.-Amer. 1: 246, plate 24. (1803) Torrey & A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 2(1): 129. (1857)
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