Flora of North America species comparison

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Eriogonum thymoides

Eriogonum brevicaule

thyme buckwheat, thyme wild buckwheat, thyme-leaf buckwheat, thyme-leaf wild buckwheat

shortstem buckwheat, shortstem wild buckwheat
Habit Subshrubs, spreading, polygamodioecious, 0.5–2(–3) × 1–3(–4) dm, tomentose to sericeous. Herbs, matted, cespitose, pulvinate, erect or spreading, sometimes scapose, (0.3–)1–5 × 1–5(–8) dm, tomentose to floccose or glabrous, grayish or greenish to green.
Stems

caudex spreading;

aerial flowering stems erect, slender, solid, not fistulose, arising at nodes of caudex branches and at distal nodes of short, nonflowering aerial branches, (0.1–)0.3–0.8(–1.2) dm, tomentose to sericeous, with a whorl of 6–10(–12), leaflike bracts ca. midlength, these similar to leaf blades, 0.3–0.8(–1) × 0.1–0.2 cm.

matted to spreading, occasionally with persistent leaf bases, up to 1/4 or more height of plant;

caudex stems matted or spreading;

aerial flowering stems spreading to erect or nearly so, slender, rarely stout, solid, not fistulose, (0.4–)0.5–2(–2.5) dm, glabrous, floccose, or sparsely to densely tomentose to lanate.
Leaves

in compact basal rosettes, fasciculate, and sheathing up stems;

petiole 0.05–0.2 cm;

blade linear to narrowly spatulate, (0.2–)0.3–1(–1.5) × 0.1–0.2(–0.3) cm, densely white-tomentose abaxially, silky-villous or sericeous adaxially, margins entire, inrolled to tightly revolute.

basal or more commonly sheathing 1–7(–15) cm up stem, 1 per node;

petiole 0.2–2(–4) cm, tomentose to floccose;

blade linear, oblanceolate, or spatulate to elliptic, (0.2–)1–10(–12) × 0.1–0.9(–1.2) cm, densely tomentose abaxially, less so to floccoseadaxially, margins plane or revolute, sometimes crenulate.
Inflorescences

capitate, 0.8–2 cm wide;

branches absent;

bracts absent immediately below involucre.

cymose, subumbellate, umbellate, or capitate, (1–)3–10(–25) × (0.7–)1–10(–15) cm;

branches dichotomous, sometimes absent, tomentose to floccose or glabrous;

bracts 3, triangular, scalelike, 1–3(–5) mm.
Peduncles

absent or erect, 0.3–3 cm, tomentose to floccose or glabrous.
Involucres

1 per node, turbinate, 3–5 × 2.5–4 mm, villous to sericeous;

teeth 6–8, erect, 0.5–1 mm.

1 per node or 3–7(–9) per cluster, turbinate to turbinate-campanulate, 1.5–4(–5) × (1–)1.5–3(–3.5) mm, tomentose to floccose or glabrous;

teeth 5, erect to spreading, 0.3–1 mm.
Flowers

4–10 mm, including 0.5–1 mm stipelike base;

perianth white to pale yellow or yellow, becoming pink to rose, villous abaxially;

tepals monomorphic, obovate;

stamens included to slightly exserted, 2–4 mm;

filaments pilose proximally.

(1–)2–4 mm;

perianth various shades of white to cream or yellow, glabrous or pubescent;

tepals connate proximal 1/4–1/3, monomorphic, lanceolate, oblong to obovate or ovate to oval;

stamens exserted, 2–4 mm;

filaments pilose basally.
Achenes

light brown, 2–2.5 mm, glabrous except for densely pubescent beak.

light brown to brown, 2–3 mm, glabrous except for roughened to papillate beak.

Eriogonum thymoides

Eriogonum brevicaule
Phenology Flowering Apr–Jul.
Habitat Sandy to gravelly, often volcanic flats, slopes, and outcrops, mixed grassland and sagebrush communities, montane conifer woodlands
Elevation (200-)600-1700 m ((700-)2000-5600 ft)
Distribution
from FNA
ID; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; ID; MT; NE; NV; SD; UT; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Eriogonum thymoides is an exquisite species concentrated in three regions of the Pacific Northwest. The first is along the eastern edge of the Cascade Range from near Wenatchee, Washington (Adams, Benton, Chelan, Douglas, Franklin, Grant, Kittitas, Klickitat, Lincoln, and Yakima counties), to near the Dalles in extreme north-central Oregon (Union County). The second is from Baker and northern Malheur counties, Oregon, to Adams, Canyon, and Washington counties, Idaho. A third series of populations is in the Mount Bennett Hills area of Gooding County, Idaho, and just over the borders in Blaine, Camas, Elmore, and Lincoln counties. Staminate plants tend to have yellow flowers that quickly fade after pollen release. Pistillate plants tend to have white to pale yellow flowers that persist and greatly elongate as the achene matures.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 8 (8 in the flora).

Eriogonum brevicaule is highly variable, and the variation has yet to be fully resolved taxonomically. The expressions recognized here will encompass the vast majority of populations. The extreme variation previously under the name var. laxifolium is now reduced with the recognition of var. bannockense (low-elevation or northern phase), var. nanum, and var. caelitum (high-elevation, southern phases).

Essentially all of the following species (28–63 below) belong to the Eriogonum brevicaule complex. Eriogonum desertorum, E. loganum, E. spathulatum, E. ostlundii, and E. artificis are allied to the complex associated with var. laxifolium, while E. natum is related to var. cottamii. Eriogonum viridulum and E. ephedroides are allied to E. brevicaule var. brevicaule as are E. contortum and E. acaule. Eriogonum brandegeei is also related, but exactly how is less certain. Allied to this complex of species are on the one hand those related to E. batemanii, and on the other all of the matted perennials belonging to the E. ochrocephalum complex. Essentially all of these species form relatively small populations on discrete edaphic sites and are well isolated one from the other. Unfortunately, a clear separation of E. brevicaule from E. desertorum, E. loganum, and E. spathulatum is not always possible.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key
1. Flowering stems and inflorescence branches glabrous
→ 2
1. Flowering stems and inflorescence branches floccose to tomentose or lanate
→ 3
2. Plants not pulvinate or cespitose, (0.8-)1.5-5 dm; inflorescences cymose, open, divided 3 times or more; widespread
var. brevicaule
2. Plants pulvinate and cespitose, 0.3-1.5(-1.8) dm; inflorescences capitate or umbellate to cymose and divided 1-2 times; Utah
var. nanum
3. Perianths pubescent; se Montana, ne Wyoming
var. canum
3. Perianths glabrous; sc Idaho, ne Nevada, Utah, sw Wyoming, not se Montana or ne Wyoming
→ 4
4. Inflorescences divided (2-)3-5 times; perianths usually yellow, rarely ochroleucous
→ 5
4. Inflorescences capitate or umbellate to cymose and divided 1-2 times; perianths ochroleucous or yellow
→ 6
5. Flowers (1-)1.5-2.5 mm; sw Wyoming
var. micranthum
5. Flowers (2.5-)3-4 mm; Utah
var. cottamii
6. Leaf blades linear to narrowly oblanceolate; inflorescences capitate or divided
→ 7
6. Leaf blades linear, oblanceolate, or narrowly oblanceolate to elliptic; inflorescences capitate. [8. Shifted to left margin.—Ed.]
→ 8
7. Leaf blades (1.5-)3-9(-12) × 0.1-0.5(-0.7) cm, tomentose abaxially, less so and grayish or occasionally greenish adaxially, margins usually revolute, occasionally plane; perianths ochroleucous or yellow; n Utah, se Idaho, below 2800 m
var. laxifolium
7. Leaf blades 0.2-4.5(-5) × (0.2-)0.3-0.6(-0.7) cm, tomentose abaxially, thinly floccose and bright green adaxially, margins plane or slightly thickened; perianths yellow; c Utah, above 2700 m
var. caelitum
8. Leaf blades and flowering stems bright green under the thinly floccose pubescence; ne Nevada and nw Utah
var. laxifolium
8. Leaf blades and flowering stems usually grayish to dull greenish under tomentum, rarely thinly floccose; se Idaho and sw Wyoming s to n Utah and ne Nevada
→ 9
9. Leaf margins crenulate; leaf blades narrowly oblanceolate to narrowly elliptic, (0.3-)0.5-1.5(-2) × 0.2-0.5(-7) cm, densely white-tomentose abaxially, floccose and greenish adaxially
var. nanum
9. Leaf margins usually plane (rarely crenulate in Wyoming); leaf blades narrowly oblanceolate to oblanceolate, (0.8-)1-4(-4.5) × (0.3-)0.4-0.8 cm, densely tomentose abaxially, tomentose to floccose and grayish to greenish adaxially
var. bannockense
Source FNA vol. 5, p. 360. FNA vol. 5, p. 264.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Oligogonum Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. brevicaule var. bannockense, E. brevicaule var. brevicaule, E. brevicaule var. caelitum, E. brevicaule var. canum, E. brevicaule var. cottamii, E. brevicaule var. laxifolium, E. brevicaule var. micranthum, E. brevicaule var. nanum
Synonyms E. sphaerocephalum subsp. minimum, E. thymoides subsp. congestum E. campanulatum subsp. brevicaule
Name authority Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 14: 9. (1856) Nuttall: Proc. Acad. Nat. Sci. Philadelphia 4: 15. (1848)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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