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Blue Mountain buckwheat, strict buckwheat

jointed buckwheat, napkinring, napkinring wild buckwheat

Habit Herbs, erect, loosely to densely matted, not scapose, 1–5 × 1–10 dm, tomentose or glabrous. Herbs, 6–15 × 0.3–0.9 dm, grayish.
Stems

spreading to erect, with or without persistent leaf bases, up to 1/4 height of plant;

caudex stems absent or spreading to matted;

aerial flowering stems erect to spreading, slender, solid, not fistulose, 1–3 dm, tomentose or glabrous.

Aerial flowering stems

5–12 dm, disarticulating into ringlike segments 3–10(–16) mm.

Leaves

basal, 1 per node;

petiole rarely twisted or curled, 1–6 cm, mostly tomentose;

blade elliptic to ovate, 0.5–2.5(–4) × (0.3–)0.5–1.5 cm, lanate, tomentose to floccose on both surfaces, sometimes sparsely tomentose to floccose and greenish or floccose to subglabrous or glabrous adaxially, margins plane.

petiole 3–8 cm, pilose;

blade oblong-ovate, 2.5–7 × 0.7–2(–3) cm, pilose on both surfaces, margins entire.

Inflorescences

umbellate-cymose to cymose, 1–20 × 3–25 cm;

branches dichotomous, tomentose to floccose or less often glabrous;

bracts 3, scalelike, triangular, 1–3 mm.

1–3 × 2–6 cm;

bracts scalelike distally, triangular, and 1–3 mm, semileaflike proximally, oblong, and 3–6 mm.

Peduncles

absent.

Involucres

1 per node, rarely 2–5 per cluster, narrowly turbinate to turbinate-campanulate, 4–6 × 1.5–5 mm, tomentose or glabrous;

teeth 5, erect, 0.5–1.3 mm.

2.5–3.5 × 3–5 mm, pilose;

teeth 0.5–2 mm.

Flowers

3–5(–6) mm;

perianth yellow or white to rose or purple, glabrous;

tepals connate proximally, dimorphic, those of outer whorl elliptic to nearly orbiculate, 2–3 × 2–3 mm, those of inner whorl oblanceolate to oblong, 3–4 × 1–2 mm;

stamens included to slightly exserted, 2–5 mm;

filaments pilose proximally.

1.5–3 mm;

tepals: those of outer whorl oblanceolate, 0.9–1.2 mm wide, those of inner whorl broadly oblanceolate to fan-shaped, 1–1.5 mm wide;

stamens 0.8–1.5 mm.

Achenes

light brown to brown, 3–3.5 mm, glabrous.

2–2.5 mm.

2n

= 40.

Eriogonum strictum

Eriogonum intrafractum

Phenology Flowering May–Oct.
Habitat Limestone washes, slopes, and cliffs, saltbush and creosote bush communities, juniper woodlands
Elevation (600-)800-1600 m ((2000-)2600-5200 ft)
Distribution
from FNA
CA; ID; MT; NV; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

Eriogonum strictum, E. niveum, and E. ovalifolium form a complex of closely related species differing in leaf, inflorescence branching, and flower features. Variety proliferum appears to be the basal entity of the complex, approaching both E. niveum and E. ovalifolium var. pansum in its pubescence and branching pattern. Also, specimens of var. proliferum are sometimes difficult to differentiate from E. nudum var. oblongifolium. Careful observation, though, will permit well-made collections to be easily distributed among the individual species. An alternative taxonomy is to reduce all of the taxa to E. ovalifolium and recognize a series of subspecies and varieties. It is possible that additional study will show that E. strictum is sufficiently distinct from its tomentose to floccose counterparts to justify recognition of E. proliferum. In that case, both var. anserinum and var. greenei would be assigned to the latter species. Or, one could follow C. L. Hitchcock et al. (1955–1969, vol. 2) and recognize subsp. strictum as distinct from subsp. proliferum, with the latter consisting of varieties proliferum, anserinum, and greenei.

Members of the Eriogonum strictum are food plants for the Bauer’s dotted-blue butterfly (Euphilotes baueri).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Rare and restricted to the Cottonwood, Funeral, Grapevine, and Panamint ranges of Death Valley National Park, Inyo County, Eriogonum intrafractum is one of the more remarkable species of the genus. The tall, simple, glabrous stem is branched only near the top, and then with only two to five inflorescence branches. The small leaves are restricted to the base of the plant. The lower part of the stem, when dried, fragments into napkinring-like segments. The rings have no evident value to the species, as they are not capable of vegetative reproduction. Some parenchyma cells at the fracture points are large enough to be seen with the naked eye, and it is likely that they act as compression points. Much as the intervertebral disks in the vertebral column provide a degree of flexibility, so too may these large parenchyma cells allow the top-heavy flowering stem to move in the wind without breaking. The flowers are numerous, each involucre so filled with more than a hundred flowers that it is torn apart by late anthesis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Inflorescence branches glabrous
var. strictum
1. Inflorescence branches tomentose to floccose
→ 2
2. Perianths yellow
var. anserinum
2. Perianths white to rose or purple
→ 3
3. Leaf blades grayish-tomentose to floccose on both surfaces, or greenish-tomentose to floccose adaxially
var. proliferum
3. Leaf blades densely whitish-lanate to tomentose on both surfaces
var. greenei
Source FNA vol. 5, p. 317. FNA vol. 5, p. 328.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Clastomyelon
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. strictum var. anserinum, E. strictum var. greenei, E. strictum var. proliferum, E. strictum var. strictum
Name authority Bentham: Trans. Linn. Soc. London 17: 414. (1836) Coville & C. V. Morton: J. Wash. Acad. Sci. 26: 305. (1936)
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