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Shockley's buckwheat, Shockley's wild buckwheat

Zion buckwheat

Habit Herbs, matted, scapose, 0.3–0.5(–0.7) × (0.5–)1–4(–20) dm, floccose to tomentose, greenish or grayish. Herbs, erect to spreading, 3–8(–10) × 0.5–1.5 dm, glabrous and glaucous, rarely tomentose, greenish.
Stems

matted, sometimes only seemingly so, with persistent leaf bases, up to 1/5 height of plant;

caudex stems matted;

aerial flowering stems absent or scapelike, erect or nearly so, slender, solid, not fistulose, (0.05–)0.1–0.3 dm, floccose to tomentose.

spreading to erect, without persistent leaf bases, up to 1/8 height of plant;

caudex stems absent;

aerial flowering stems erect to spreading, slender to stout, solid or hollow, often fistulose, 1–2.5(–3) dm, glabrous, rarely tomentose.

Leaves

basal, fasciculate in terminal tufts;

petiole 0.2–0.5 cm, tomentose to floccose;

blade oblanceolate to elliptic or spatulate, (0.2–)0.3–0.8(–1.2) × 0.2–0.4(–0.6) cm, tomentose to floccose, margins plane or slightly thickened.

basal, 1 per node;

petiole 3–6(–10) cm, tomentose to floccose;

blade elliptic or oblong-ovate to ovate, 2–4.5(–6) × 1.5–2.5(–5) cm, lanate to densely tomentose abaxially, thinly floccose or glabrous and green adaxially, margins plane.

Inflorescences

capitate, 0.8–2 cm;

branches absent;

bracts 3–5, scalelike, linear to linear-lanceolate, 1.5–4 × 0.6–1 mm.

narrowly virgate or racemose with involucres racemosely disposed throughout or at tips, 20–50(–60) × 5–25 cm, glabrous, rarely tomentose;

branches dichotomous, upper secondaries suppressed and bearing (8–)10–15(–20) racemosely arranged involucres;

bracts 3, scalelike, triangular, and 2–7 mm, or leaflike, elliptic, and 10–30 × 4–15 mm.

Peduncles

absent.

absent.

Involucres

2–4(–6) per cluster, campanulate, (2–)2.5–5(–6) × 3–6(–7) mm, rigid, tomentose;

teeth 5–10, erect to spreading, (0.5–)1–3 mm.

1 per node, turbinate to turbinate-campanulate, 1.5–3 × 1.5–2.5 mm, tomentose;

teeth 5, erect, 0.2–0.4 mm.

Flowers

2.5–4 mm;

perianth white to rose or yellow, densely pilose;

tepals connate proximally, monomorphic, oblong to obovate;

stamens exserted, 2.5–5 mm;

filaments subglabrous or sparsely pilose proximally.

2–6(–7) mm;

perianth white to yellow or bright red, glabrous;

tepals connate proximal 1/4 their length, monomorphic, oblong;

stamens exserted, 2–5 mm;

filaments pilose proximally.

Achenes

light brown to brown, 2.5–3 mm, tomentose.

light brown, (3–)4–5(–6) mm, glabrous.

Eriogonum shockleyi

Eriogonum zionis

Phenology Flowering May–Aug.
Habitat Gravelly or clayey (rarely sandy) flats, washes, and slopes, saltbush, blackbrush, and sagebrush communities, pinyon-juniper woodlands
Elevation (800-)1200-2600 m ((2600-)3900-8500 ft)
Distribution
from FNA
AZ; CA; CO; ID; NM; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; UT
[BONAP county map]
Discussion

Eriogonum shockleyi is widely distributed in northern Arizona, east-central California, western Colorado, southern Idaho, northwestern New Mexico, Nevada, and Utah. On the Colorado Plateau, it has oblanceolate to spatulate leaf blades usually 0.3–1.2 × 0.3–0.6 cm, scapes 1–3 cm, involucres with long (2–3.5 mm) often spreading teeth, and flowers 3–4 mm. These plants have been distinguished as var. longilobum. The typical Great Basin expression has elliptic leaf blades 0.3–0.6 × 0.3–0.5 cm, scapes 0.5–2 cm, involucres with short (0.5–2 mm) erect teeth, and flowers 2.5–4 mm. Low, compact, hummock-like plants in southwestern Idaho with elliptic leaf blades 0.1–0.3(–0.35) × 0.1–0.15 cm, flowering stems absent or up to 0.5 cm, and involucres 2.5–3 mm with teeth 0.8–1 mm have been termed var. packardiae. These morphologic differences have been shown to be genetically insignificant, however (J. F. Smith and T. A. Bateman 2002). Plants on moving sand dunes at the southeast end of Baking Power Flat in Lincoln County, Nevada, can be up to 2 m across. Further studies may indicate that the various geographic expressions deserve taxonomic recognition.

The Great Basin expression is the food plant of the Bernardino dotted-blue butterfly (Euphilotes bernardino). Members of the species are occasionally found in cultivation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Perianths white to yellow
var. zionis
1. Perianths bright red
var. coccineum
Source FNA vol. 5, p. 286. FNA vol. 5, p. 297.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii
Subordinate taxa
E. zionis var. coccineum, E. zionis var. zionis
Synonyms E. pulvinatum, E. shockleyi subsp. candidum, E. shockleyi subsp. longilobum, E. shockleyi var. longilobum, E. shockleyi var. packardiae, E. villiflorum var. candidum E. racemosum var. zionis
Name authority S. Watson: Proc. Amer. Acad. Arts 18: 194. (1883) J. T. Howell: Leafl. W. Bot. 2: 253. (1940)
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