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Shockley's buckwheat, Shockley's wild buckwheat
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Twisselmann's buckwheat, Twisselmann's wild buckwheat
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| Herbs, matted, scapose, 0.3–0.5(–0.7) × (0.5–)1–4(–20) dm, floccose to tomentose, greenish or grayish. |
Herbs, matted, occasionally polygamodioecious, (0.2–)0.5–1.5 × 3–4 dm, tomentose. |
matted, sometimes only seemingly so, with persistent leaf bases, up to 1/5 height of plant; caudex stems matted; aerial flowering stems absent or scapelike, erect or nearly so, slender, solid, not fistulose, (0.05–)0.1–0.3 dm, floccose to tomentose. |
caudex spreading; aerial flowering stems erect or nearly so, slender, solid, not fistulose, arising at nodes of caudex branches and at distal nodes of short, nonflowering aerial branches, (0.2–)0.5–1.2 dm, tomentose, with a whorl of 4–8 leaflike bracts ca. midlength, similar to leaf blades, 0.4–0.8 × 0.1–0.3 cm. |
basal, fasciculate in terminal tufts; petiole 0.2–0.5 cm, tomentose to floccose; blade oblanceolate to elliptic or spatulate, (0.2–)0.3–0.8(–1.2) × 0.2–0.4(–0.6) cm, tomentose to floccose, margins plane or slightly thickened. |
in loose to congested basal rosettes; petiole 0.2–0.4 cm, thinly tomentose; blade oblanceolate to elliptic, 0.5–1 × (0.2–)0.4–0.6 cm, white- or grayish-tomentose abaxially, thinly tomentose to nearly glabrous and dull green adaxially. |
capitate, 0.8–2 cm; branches absent; bracts 3–5, scalelike, linear to linear-lanceolate, 1.5–4 × 0.6–1 mm. |
capitate, 0.7–1.5 cm wide; branches absent; bracts absent immediately below involucre. |
absent. |
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2–4(–6) per cluster, campanulate, (2–)2.5–5(–6) × 3–6(–7) mm, rigid, tomentose; teeth 5–10, erect to spreading, (0.5–)1–3 mm. |
1 per node, campanulate, (2.5–)4–5 × 4–6 mm; teeth 6–9, lobelike, strongly reflexed, (2–)3–5 mm. |
2.5–4 mm; perianth white to rose or yellow, densely pilose; tepals connate proximally, monomorphic, oblong to obovate; stamens exserted, 2.5–5 mm; filaments subglabrous or sparsely pilose proximally. |
(4–)5–6 mm, including 1–1.3 mm stipelike base; perianth pale yellow, villous abaxially; tepals slightly dimorphic, those of outer whorl obtuse, 3–5 × 3–3.5 mm, those of inner whorl broadly oblanceolate, 5–6 × 2.5–3 mm; stamens exserted, 4–5 mm; filaments pilose proximally. |
light brown to brown, 2.5–3 mm, tomentose. |
light brown, 5–5.5 mm, glabrous except for sparsely pubescent beak. |
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| Flowering May–Aug. |
Flowering Jun–Sep. |
| Gravelly or clayey (rarely sandy) flats, washes, and slopes, saltbush, blackbrush, and sagebrush communities, pinyon-juniper woodlands |
Rocky outcrops, montane conifer woodlands |
| (800-)1200-2600 m ((2600-)3900-8500 ft) |
2300-2600 m (7500-8500 ft) |
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AZ; CA; CO; ID; NM; NV; UT
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CA |
Eriogonum shockleyi is widely distributed in northern Arizona, east-central California, western Colorado, southern Idaho, northwestern New Mexico, Nevada, and Utah. On the Colorado Plateau, it has oblanceolate to spatulate leaf blades usually 0.3–1.2 × 0.3–0.6 cm, scapes 1–3 cm, involucres with long (2–3.5 mm) often spreading teeth, and flowers 3–4 mm. These plants have been distinguished as var. longilobum. The typical Great Basin expression has elliptic leaf blades 0.3–0.6 × 0.3–0.5 cm, scapes 0.5–2 cm, involucres with short (0.5–2 mm) erect teeth, and flowers 2.5–4 mm. Low, compact, hummock-like plants in southwestern Idaho with elliptic leaf blades 0.1–0.3(–0.35) × 0.1–0.15 cm, flowering stems absent or up to 0.5 cm, and involucres 2.5–3 mm with teeth 0.8–1 mm have been termed var. packardiae. These morphologic differences have been shown to be genetically insignificant, however (J. F. Smith and T. A. Bateman 2002). Plants on moving sand dunes at the southeast end of Baking Power Flat in Lincoln County, Nevada, can be up to 2 m across. Further studies may indicate that the various geographic expressions deserve taxonomic recognition. The Great Basin expression is the food plant of the Bernardino dotted-blue butterfly (Euphilotes bernardino). Members of the species are occasionally found in cultivation. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Eriogonum twisselmannii is a rare species of the southern Sierra Nevada and is known only from Slate Mountain and The Needles in Tulare County. It is worthy of being on a “watch” list of rare plants in California, but because of its remote location, it is not threatened. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 5, p. 286. |
FNA vol. 5, p. 364. |
| Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Oligogonum |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
| E. pulvinatum, E. shockleyi subsp. candidum, E. shockleyi subsp. longilobum, E. shockleyi var. longilobum, E. shockleyi var. packardiae, E. villiflorum var. candidum |
E. douglasii var. twisselmannii |
| S. Watson: Proc. Amer. Acad. Arts 18: 194. (1883) |
(J. T. Howell) Reveal: in P. A. Munz, Suppl. Calif. Fl. 40. (1968) |
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