Eriogonum parvifolium |
Eriogonum strictum |
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dune wild buckwheat, sea cliff buckwheat, seacliff wild buckwheat |
Blue Mountain buckwheat, strict buckwheat |
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Habit | Shrubs, matted to spreading or rounded, 3–10 × 5–20(–25) dm, thinly tomentose or glabrous, greenish. | Herbs, erect, loosely to densely matted, not scapose, 1–5 × 1–10 dm, tomentose or glabrous. | ||||||||||||
Stems | spreading, sometimes matted, often with persistent leaf bases, up to 1/2 or more height of plant; caudex stems absent or matted; aerial flowering stems prostrate, spreading, or erect, slender, solid, not fistulose, 0.2–1 dm, thinly tomentose or glabrous. |
spreading to erect, with or without persistent leaf bases, up to 1/4 height of plant; caudex stems absent or spreading to matted; aerial flowering stems erect to spreading, slender, solid, not fistulose, 1–3 dm, tomentose or glabrous. |
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Leaves | cauline, fasciculate, infrequently 1 per node; petiole 0.1–0.7 cm, floccose; blade lanceolate to round, 0.5–3 × 0.3–0.8(–1.2) cm, lanate to tomentose abaxially, mostly glabrous and olive green to green adaxially. |
basal, 1 per node; petiole rarely twisted or curled, 1–6 cm, mostly tomentose; blade elliptic to ovate, 0.5–2.5(–4) × (0.3–)0.5–1.5 cm, lanate, tomentose to floccose on both surfaces, sometimes sparsely tomentose to floccose and greenish or floccose to subglabrous or glabrous adaxially, margins plane. |
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Inflorescences | capitate to cymose, 20–30 × 2–10 cm; branches dichotomous, thinly tomentose or glabrous; bracts 3, scalelike, triangular, and 1–2 mm, or leaflike, usually elliptic, and 5–20 × 2–10 mm. |
umbellate-cymose to cymose, 1–20 × 3–25 cm; branches dichotomous, tomentose to floccose or less often glabrous; bracts 3, scalelike, triangular, 1–3 mm. |
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Peduncles | absent. |
absent. |
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Involucres | 2–7 per cluster, turbinate-campanulate, (2.5–)3–4 × 2–3.5 mm, floccose to glabrate; teeth 5, erect, 0.5–0.9 mm. |
1 per node, rarely 2–5 per cluster, narrowly turbinate to turbinate-campanulate, 4–6 × 1.5–5 mm, tomentose or glabrous; teeth 5, erect, 0.5–1.3 mm. |
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Flowers | 2.5–3 mm; perianth white to pinkish or greenish yellow, glabrous; tepals connate proximally, monomorphic, obovate; stamens exserted, 2.5–3.5 mm; filaments pilose proximally. |
3–5(–6) mm; perianth yellow or white to rose or purple, glabrous; tepals connate proximally, dimorphic, those of outer whorl elliptic to nearly orbiculate, 2–3 × 2–3 mm, those of inner whorl oblanceolate to oblong, 3–4 × 1–2 mm; stamens included to slightly exserted, 2–5 mm; filaments pilose proximally. |
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Achenes | brown, 2.5–3 mm, glabrous. |
light brown to brown, 3–3.5 mm, glabrous. |
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2n | = 40. |
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Eriogonum parvifolium |
Eriogonum strictum |
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Phenology | Flowering year-round. | |||||||||||||
Habitat | Sandy beaches, dunes, and bluffs or sandy to gravelly inland slopes and flats, coastal grassland and chaparral communities, oak and pine woodlands | |||||||||||||
Elevation | 0-300 m (0-1000 ft) | |||||||||||||
Distribution |
CA
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CA; ID; MT; NV; OR; WA
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Discussion | The native range of Eriogonum parvifolium is restricted to coastal and near-coastal areas (Los Angeles, Monterey, Orange, San Diego, San Luis Obispo, Santa Barbara, and Ventura counties). The coastal expression (var. parvifolium) has thickened leaf blades (0.5–1.5 × 0.3–0.8 cm) and simple or dichotomous inflorescences of compact clusters of involucres containing white to rose flowers. Highly compact and dense mat-forming plants on rocky bluffs immediately next to the ocean were named var. crassifolium; those with yellow flowers were named var. lucidum. The inland form with thin leaf blades (1.5–3 × 0.3–0.8 cm) and highly-branched, cymose, white-flowered inflorescences is perhaps worthy of continued recognition as var. paynei, although there is no sharp distinction between the extremes. Several expressions of the seacliff wild buckwheat are in cultivation, and unfortunately the California Department of Transportation is using the species in roadside plantings, with the result that it is now established in Santa Clara County. Every effort should be made to halt its introduction beyond its native range. The species is the food plant for two federally endangered butterflies, the El Segundo dotted-blue (Euphilotes battoides allyni), near Los Angeles, and Smith’s dotted-blue (Euphilotes enoptes smithi), near Monterey. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 4 (4 in the flora). Eriogonum strictum, E. niveum, and E. ovalifolium form a complex of closely related species differing in leaf, inflorescence branching, and flower features. Variety proliferum appears to be the basal entity of the complex, approaching both E. niveum and E. ovalifolium var. pansum in its pubescence and branching pattern. Also, specimens of var. proliferum are sometimes difficult to differentiate from E. nudum var. oblongifolium. Careful observation, though, will permit well-made collections to be easily distributed among the individual species. An alternative taxonomy is to reduce all of the taxa to E. ovalifolium and recognize a series of subspecies and varieties. It is possible that additional study will show that E. strictum is sufficiently distinct from its tomentose to floccose counterparts to justify recognition of E. proliferum. In that case, both var. anserinum and var. greenei would be assigned to the latter species. Or, one could follow C. L. Hitchcock et al. (1955–1969, vol. 2) and recognize subsp. strictum as distinct from subsp. proliferum, with the latter consisting of varieties proliferum, anserinum, and greenei. Members of the Eriogonum strictum are food plants for the Bauer’s dotted-blue butterfly (Euphilotes baueri). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 300. | FNA vol. 5, p. 317. | ||||||||||||
Parent taxa | Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla | Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla | ||||||||||||
Sibling taxa | ||||||||||||||
Subordinate taxa | ||||||||||||||
Synonyms | E. parvifolium var. crassifolium, E. parvifolium subsp. lucidum, E. parvifolium var. lucidum, E. parvifolium subsp. paynei, E. parvifolium var. paynei | |||||||||||||
Name authority | Smith: in A. Rees, Cycl. 13(2): Eriogonum no. 2. (1809) | Bentham: Trans. Linn. Soc. London 17: 414. (1836) | ||||||||||||
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