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dune wild buckwheat, sea cliff buckwheat, seacliff wild buckwheat

ladder buckwheat, ladder wild buckwheat

Habit Shrubs, matted to spreading or rounded, 3–10 × 5–20(–25) dm, thinly tomentose or glabrous, greenish. Herbs, erect, annual, (0.5–)3–6(–10) dm, glabrous, glaucous, grayish.
Stems

spreading, sometimes matted, often with persistent leaf bases, up to 1/2 or more height of plant;

caudex stems absent or matted;

aerial flowering stems prostrate, spreading, or erect, slender, solid, not fistulose, 0.2–1 dm, thinly tomentose or glabrous.

caudex absent;

aerial flowering stems erect, solid, not fistulose, (0.5–)1–2 dm, glabrous.

Leaves

cauline, fasciculate, infrequently 1 per node;

petiole 0.1–0.7 cm, floccose;

blade lanceolate to round, 0.5–3 × 0.3–0.8(–1.2) cm, lanate to tomentose abaxially, mostly glabrous and olive green to green adaxially.

basal;

petiole 1–10 cm, tomentose;

blade sub-cordate to orbiculate, (1.5–)2–5(–8) × (1.5–)2–5(–8) cm, densely white-tomentose abaxially, floccose to subglabrous and green adaxially, margins often wavy.

Inflorescences

capitate to cymose, 20–30 × 2–10 cm;

branches dichotomous, thinly tomentose or glabrous;

bracts 3, scalelike, triangular, and 1–2 mm, or leaflike, usually elliptic, and 5–20 × 2–10 mm.

cymose, spreading when immature, quickly becoming narrowly erect and strict with whiplike branches, (5–)10–50(–80) × 10–50 cm;

branches glabrous, glaucous;

bracts 3, scalelike, 1–1.5(–2) × 0.4–1 mm.

Peduncles

absent.

absent or erect, straight, slender, 0.1–0.2 cm, glabrous.

Involucres

2–7 per cluster, turbinate-campanulate, (2.5–)3–4 × 2–3.5 mm, floccose to glabrate;

teeth 5, erect, 0.5–0.9 mm.

turbinate, 2–2.5(–3) × 1.5–2.5 mm, glabrous;

teeth 5, erect, 0.5–1.5 mm.

Flowers

2.5–3 mm;

perianth white to pinkish or greenish yellow, glabrous;

tepals connate proximally, monomorphic, obovate;

stamens exserted, 2.5–3.5 mm;

filaments pilose proximally.

1.5–2 mm;

perianth white with green or reddish midribs, becoming pinkish, glabrous;

tepals dimorphic, those of outer whorl oblong, those of inner whorl lanceolate;

stamens exserted, 1.5–2.5 mm;

filaments pilose proximally.

Achenes

brown, 2.5–3 mm, glabrous.

dark brown to blackish, 3-gonous, 2–2.5 mm, glabrous.

2n

= 40.

= 40.

Eriogonum parvifolium

Eriogonum exaltatum

Phenology Flowering year-round. Flowering May–Oct.
Habitat Sandy beaches, dunes, and bluffs or sandy to gravelly inland slopes and flats, coastal grassland and chaparral communities, oak and pine woodlands Sandy to gravelly flats and slopes, saltbush, creosote bush, greasewood, blackbrush, and mesquite communities, rarely in pinyon-juniper woodlands
Elevation 0-300 m (0-1000 ft) 500-1400 m (1600-4600 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NV; UT
[BONAP county map]
Discussion

The native range of Eriogonum parvifolium is restricted to coastal and near-coastal areas (Los Angeles, Monterey, Orange, San Diego, San Luis Obispo, Santa Barbara, and Ventura counties). The coastal expression (var. parvifolium) has thickened leaf blades (0.5–1.5 × 0.3–0.8 cm) and simple or dichotomous inflorescences of compact clusters of involucres containing white to rose flowers. Highly compact and dense mat-forming plants on rocky bluffs immediately next to the ocean were named var. crassifolium; those with yellow flowers were named var. lucidum. The inland form with thin leaf blades (1.5–3 × 0.3–0.8 cm) and highly-branched, cymose, white-flowered inflorescences is perhaps worthy of continued recognition as var. paynei, although there is no sharp distinction between the extremes. Several expressions of the seacliff wild buckwheat are in cultivation, and unfortunately the California Department of Transportation is using the species in roadside plantings, with the result that it is now established in Santa Clara County. Every effort should be made to halt its introduction beyond its native range.

The species is the food plant for two federally endangered butterflies, the El Segundo dotted-blue (Euphilotes battoides allyni), near Los Angeles, and Smith’s dotted-blue (Euphilotes enoptes smithi), near Monterey.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eriogonum insigne has been a troublesome taxon.  The type, collected by E. Palmer in 1876, is a curious specimen from near Paragonah in Iron County, Utah.  That expression has not been recollected and, after another frustrating review of the type material, it is now referred to E. deflexum, along with other odd specimens from southern Nevada and California that have always been difficult to assign.  As a result, the tall, upright plants with long, whiplike branches of northwestern Arizona (Mohave County), southern Nevada (northeastern Clark and southern Lincoln counties), and southwestern Utah (southwestern Washington County), are now recognized under the name E. exaltatum.  Some specimens remain problematic.  Immature plants of E. exaltatum (Goodding 2302, GH, MIN, MO, NY, RM, UC) resemble E. bifurcatum. Specimens from Eureka Valley (DeDecker 4741, NY) and western Pahrump Valley (Abrams 14248, DS, GH, NY) in Inyo County, California, appear to belong to E. exaltatum.

Like Eriogonum exaltatum, some populations of E. deflexum var. deflexum have nearly erect involucres.  The plants from the Buried Hills (Nye and Clark counties, Nevada) are particularly odd.  Single plants from a few collections in Kane County, Utah, have sessile, seemingly erect involucres in the forks of inflorescence branches.  None of these plants has the characteristic long, whiplike branches of E. exaltatum, and they are not included in E. deflexum var. deflexum. The type of E. deflexum var. rectum from San Bernardino County, California (Reveal & Broome 6385, CAS, NY, US, etc.), has individual specimens with both deflexed and somewhat erect involucres, along with whiplike branches.  Collections from Imperial and San Diego counties, previously associated with what is here termed E. exaltatum, are now considered specimens of E. deflexum.  Clearly, more work is required to understand these species fully.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 300. FNA vol. 5, p. 397.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Synonyms E. parvifolium var. crassifolium, E. parvifolium subsp. lucidum, E. parvifolium var. lucidum, E. parvifolium subsp. paynei, E. parvifolium var. paynei
Name authority Smith: in A. Rees, Cycl. 13(2): Eriogonum no. 2. (1809) M. E. Jones: Contr. W. Bot. 15: 61. (1929)
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