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dune wild buckwheat, sea cliff buckwheat, seacliff wild buckwheat
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spreading buckwheat, spreading wild buckwheat
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| Shrubs, matted to spreading or rounded, 3–10 × 5–20(–25) dm, thinly tomentose or glabrous, greenish. |
Shrubs, spreading, not scapose, (1.5–)2–5(–7) × 5–15 dm, grayish- to reddish-brown-tomentose to floccose and gray or, rarely, thinly floccose and greenish. |
spreading, sometimes matted, often with persistent leaf bases, up to 1/2 or more height of plant; caudex stems absent or matted; aerial flowering stems prostrate, spreading, or erect, slender, solid, not fistulose, 0.2–1 dm, thinly tomentose or glabrous. |
spreading to erect, typically without persistent leaf bases, up to 1/2 height of plant; caudex stems absent; aerial flowering stems erect to spreading, slender, solid, not fistulose, 0.3–0.8 dm, floccose or glabrous. |
cauline, fasciculate, infrequently 1 per node; petiole 0.1–0.7 cm, floccose; blade lanceolate to round, 0.5–3 × 0.3–0.8(–1.2) cm, lanate to tomentose abaxially, mostly glabrous and olive green to green adaxially. |
cauline, 1 per node; petiole 0.2–0.7 cm, tomentose to floccose; blade oblanceolate to oblong or obovate, (1–)1.5–3 × (0.2–)0.3–0.7 cm, densely white-tomentose abaxially, white-floccose to glabrate or green and glabrous adaxially, margins plane. |
capitate to cymose, 20–30 × 2–10 cm; branches dichotomous, thinly tomentose or glabrous; bracts 3, scalelike, triangular, and 1–2 mm, or leaflike, usually elliptic, and 5–20 × 2–10 mm. |
cymose, 10–30(–40) × 10–40 cm; branches dichotomous, white-floccose to glabrate or subglabrous; bracts 3, scalelike, triangular, 0.5–2(–5) mm. |
absent. |
absent or mostly erect, slender, 0.3–2.5 cm, floccose. |
2–7 per cluster, turbinate-campanulate, (2.5–)3–4 × 2–3.5 mm, floccose to glabrate; teeth 5, erect, 0.5–0.9 mm. |
1 per node, turbinate, 1.5–2.5(–3) × 1–2 mm, tomentose to floccose; teeth 5, erect, 0.3–0.6 mm. |
2.5–3 mm; perianth white to pinkish or greenish yellow, glabrous; tepals connate proximally, monomorphic, obovate; stamens exserted, 2.5–3.5 mm; filaments pilose proximally. |
2–4 mm; perianth yellow, glabrous; tepals connate proximal 1/4, essentially monomorphic, elliptic to obovate; stamens mostly exserted, 2–4.5 mm; filaments sparsely pilose proximally. |
brown, 2.5–3 mm, glabrous. |
brown, 2–2.5 mm, glabrous. |
= 40. |
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| Flowering year-round. |
Flowering Jun–Sep. |
| Sandy beaches, dunes, and bluffs or sandy to gravelly inland slopes and flats, coastal grassland and chaparral communities, oak and pine woodlands |
Sandy to rocky slopes and flats, mixed grassland and sagebrush communities, juniper and montane conifer woodlands |
| 0-300 m (0-1000 ft) |
1200-2500 m (3900-8200 ft) |
|
CA
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CO; MT; NE; NM; SD; WY
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The native range of Eriogonum parvifolium is restricted to coastal and near-coastal areas (Los Angeles, Monterey, Orange, San Diego, San Luis Obispo, Santa Barbara, and Ventura counties). The coastal expression (var. parvifolium) has thickened leaf blades (0.5–1.5 × 0.3–0.8 cm) and simple or dichotomous inflorescences of compact clusters of involucres containing white to rose flowers. Highly compact and dense mat-forming plants on rocky bluffs immediately next to the ocean were named var. crassifolium; those with yellow flowers were named var. lucidum. The inland form with thin leaf blades (1.5–3 × 0.3–0.8 cm) and highly-branched, cymose, white-flowered inflorescences is perhaps worthy of continued recognition as var. paynei, although there is no sharp distinction between the extremes. Several expressions of the seacliff wild buckwheat are in cultivation, and unfortunately the California Department of Transportation is using the species in roadside plantings, with the result that it is now established in Santa Clara County. Every effort should be made to halt its introduction beyond its native range. The species is the food plant for two federally endangered butterflies, the El Segundo dotted-blue (Euphilotes battoides allyni), near Los Angeles, and Smith’s dotted-blue (Euphilotes enoptes smithi), near Monterey. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eriogonum effusum is rather common on the northern Great Plains and along the eastern slope of the Rocky Mountains in central and eastern Colorado, southeastern Montana, western Nebraska, northern New Mexico, southwestern South Dakota, and southeastern Wyoming. Some specimens from Chaffee County, Colorado, are thinly floccose and greenish (Atwood & Welsh 29689, BRY) and thus similar to E. leptocladon. A collection from Pyramid Lake, Washoe County, Nevada (Frandsen & Brown 182, NESH) is clearly mislabeled. A roadside collection of E. effusum gathered near Little America, Sweetwater County, Wyoming, in 1961 (G. Mason 4025, ASU) was an introduction that has not persisted. The spreading wild buckwheat is occasionally merged with Eriogonum microthecum even though the two are morphologically distinct and their ranges do not overlap. Plants in New Mexico are sometimes difficult to distinguish from the related E. leptocladon var. ramosissimum. The species is the food plant for the Rita dotted-blue butterfly (Euphilotes rita). A hybrid between Eriogonum effusum and E. pauciflorum has been named E. ×nebraskense Rydberg [E. multiceps Nees subsp. nebraskense (Rydberg) S. Stokes; E. pauciflorum Pursh var. nebraskense (Rydberg) Reveal]. The hybrid is known from Weld County, Colorado; Cheyenne and Kimball counties, Nebraska; and Converse and Platte counties, Wyoming. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 5, p. 300. |
FNA vol. 5, p. 248. |
| Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
| E. parvifolium var. crassifolium, E. parvifolium subsp. lucidum, E. parvifolium var. lucidum, E. parvifolium subsp. paynei, E. parvifolium var. paynei |
E. microthecum var. effusum |
| Smith: in A. Rees, Cycl. 13(2): Eriogonum no. 2. (1809) |
Nuttall: Proc. Acad. Nat. Sci. Philadelphia 4: 15. (1848) |
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CA
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