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dune wild buckwheat, sea cliff buckwheat, seacliff wild buckwheat

Cithara buckwheat, Cithara wild buckwheat

Habit Shrubs, matted to spreading or rounded, 3–10 × 5–20(–25) dm, thinly tomentose or glabrous, greenish. Herbs, spreading to erect, 2–3(–5) dm, glabrous or tomentose, greenish or reddish.
Stems

spreading, sometimes matted, often with persistent leaf bases, up to 1/2 or more height of plant;

caudex stems absent or matted;

aerial flowering stems prostrate, spreading, or erect, slender, solid, not fistulose, 0.2–1 dm, thinly tomentose or glabrous.

aerial flowering stems erect, 0.5–1 dm, glabrous or tomentose.

Leaves

cauline, fasciculate, infrequently 1 per node;

petiole 0.1–0.7 cm, floccose;

blade lanceolate to round, 0.5–3 × 0.3–0.8(–1.2) cm, lanate to tomentose abaxially, mostly glabrous and olive green to green adaxially.

basal and sometimes cauline;

basal: petiole 1–5 cm, tomentose to floccose, often slightly winged, blade oblanceolate or elliptic to ovate or nearly rounded, 1–2 × 0.5–1.5(–2) cm, tomentose abaxially, floccose to glabrate and grayish to greenish adaxially;

cauline: petiole 0.5–2 cm, floccose, blade elliptic, 0.3–1 cm × 0.2–0.7 cm, similar to basal blade.

Inflorescences

capitate to cymose, 20–30 × 2–10 cm;

branches dichotomous, thinly tomentose or glabrous;

bracts 3, scalelike, triangular, and 1–2 mm, or leaflike, usually elliptic, and 5–20 × 2–10 mm.

cymose, distally uniparous due to suppression of secondary branches, open, 5–25 × 5–25 cm;

branches usually inwardly curved, glabrous or tomentose;

bracts 0.5–2.5 × 0.5–2 mm.

Peduncles

absent.

absent.

Involucres

2–7 per cluster, turbinate-campanulate, (2.5–)3–4 × 2–3.5 mm, floccose to glabrate;

teeth 5, erect, 0.5–0.9 mm.

somewhat appressed to branches, turbinate, 2.5–3 × 1.5–2 mm, glabrous;

teeth 5, erect, 0.2–0.4 mm.

Flowers

2.5–3 mm;

perianth white to pinkish or greenish yellow, glabrous;

tepals connate proximally, monomorphic, obovate;

stamens exserted, 2.5–3.5 mm;

filaments pilose proximally.

1.5–2 mm;

perianth white to rose, glabrous;

tepals monomorphic, oblong-obovate;

stamens included, 1–1.5 mm;

filaments pilose proximally.

Achenes

brown, 2.5–3 mm, glabrous.

brown, 3-gonous, 1–1.5 mm.

2n

= 40.

Eriogonum parvifolium

Eriogonum cithariforme

Phenology Flowering year-round.
Habitat Sandy beaches, dunes, and bluffs or sandy to gravelly inland slopes and flats, coastal grassland and chaparral communities, oak and pine woodlands
Elevation 0-300 m (0-1000 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

The native range of Eriogonum parvifolium is restricted to coastal and near-coastal areas (Los Angeles, Monterey, Orange, San Diego, San Luis Obispo, Santa Barbara, and Ventura counties). The coastal expression (var. parvifolium) has thickened leaf blades (0.5–1.5 × 0.3–0.8 cm) and simple or dichotomous inflorescences of compact clusters of involucres containing white to rose flowers. Highly compact and dense mat-forming plants on rocky bluffs immediately next to the ocean were named var. crassifolium; those with yellow flowers were named var. lucidum. The inland form with thin leaf blades (1.5–3 × 0.3–0.8 cm) and highly-branched, cymose, white-flowered inflorescences is perhaps worthy of continued recognition as var. paynei, although there is no sharp distinction between the extremes. Several expressions of the seacliff wild buckwheat are in cultivation, and unfortunately the California Department of Transportation is using the species in roadside plantings, with the result that it is now established in Santa Clara County. Every effort should be made to halt its introduction beyond its native range.

The species is the food plant for two federally endangered butterflies, the El Segundo dotted-blue (Euphilotes battoides allyni), near Los Angeles, and Smith’s dotted-blue (Euphilotes enoptes smithi), near Monterey.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

The graceful, upwardly curved branch segments of Eriogonum cithariforme generally are distinctive in older plants, although an occasional individual of E. davidsonii may have this feature. A distinction between E. cithariforme and E. roseum also is troublesome, as in young material the curved branch segments are not always obvious.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Flowering stems and inflorescence branches tomentose
var. cithariforme
1. Flowering stems and inflorescence branches glabrous
var. agninum
Source FNA vol. 5, p. 300. FNA vol. 5, p. 424.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Oregonium
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. cithariforme var. agninum, E. cithariforme var. cithariforme
Synonyms E. parvifolium var. crassifolium, E. parvifolium subsp. lucidum, E. parvifolium var. lucidum, E. parvifolium subsp. paynei, E. parvifolium var. paynei E. gracile var. cithariforme, E. vimineum var. cithariforme
Name authority Smith: in A. Rees, Cycl. 13(2): Eriogonum no. 2. (1809) S. Watson: Proc. Amer. Acad. Arts 23: 266. (1888)
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