dune wild buckwheat, sea cliff buckwheat, seacliff wild buckwheat
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shortstem buckwheat, shortstem wild buckwheat
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Shrubs, matted to spreading or rounded, 3–10 × 5–20(–25) dm, thinly tomentose or glabrous, greenish. |
Herbs, matted, cespitose, pulvinate, erect or spreading, sometimes scapose, (0.3–)1–5 × 1–5(–8) dm, tomentose to floccose or glabrous, grayish or greenish to green. |
spreading, sometimes matted, often with persistent leaf bases, up to 1/2 or more height of plant; caudex stems absent or matted; aerial flowering stems prostrate, spreading, or erect, slender, solid, not fistulose, 0.2–1 dm, thinly tomentose or glabrous. |
matted to spreading, occasionally with persistent leaf bases, up to 1/4 or more height of plant; caudex stems matted or spreading; aerial flowering stems spreading to erect or nearly so, slender, rarely stout, solid, not fistulose, (0.4–)0.5–2(–2.5) dm, glabrous, floccose, or sparsely to densely tomentose to lanate. |
cauline, fasciculate, infrequently 1 per node; petiole 0.1–0.7 cm, floccose; blade lanceolate to round, 0.5–3 × 0.3–0.8(–1.2) cm, lanate to tomentose abaxially, mostly glabrous and olive green to green adaxially. |
basal or more commonly sheathing 1–7(–15) cm up stem, 1 per node; petiole 0.2–2(–4) cm, tomentose to floccose; blade linear, oblanceolate, or spatulate to elliptic, (0.2–)1–10(–12) × 0.1–0.9(–1.2) cm, densely tomentose abaxially, less so to floccoseadaxially, margins plane or revolute, sometimes crenulate. |
capitate to cymose, 20–30 × 2–10 cm; branches dichotomous, thinly tomentose or glabrous; bracts 3, scalelike, triangular, and 1–2 mm, or leaflike, usually elliptic, and 5–20 × 2–10 mm. |
cymose, subumbellate, umbellate, or capitate, (1–)3–10(–25) × (0.7–)1–10(–15) cm; branches dichotomous, sometimes absent, tomentose to floccose or glabrous; bracts 3, triangular, scalelike, 1–3(–5) mm. |
absent. |
absent or erect, 0.3–3 cm, tomentose to floccose or glabrous. |
2–7 per cluster, turbinate-campanulate, (2.5–)3–4 × 2–3.5 mm, floccose to glabrate; teeth 5, erect, 0.5–0.9 mm. |
1 per node or 3–7(–9) per cluster, turbinate to turbinate-campanulate, 1.5–4(–5) × (1–)1.5–3(–3.5) mm, tomentose to floccose or glabrous; teeth 5, erect to spreading, 0.3–1 mm. |
2.5–3 mm; perianth white to pinkish or greenish yellow, glabrous; tepals connate proximally, monomorphic, obovate; stamens exserted, 2.5–3.5 mm; filaments pilose proximally. |
(1–)2–4 mm; perianth various shades of white to cream or yellow, glabrous or pubescent; tepals connate proximal 1/4–1/3, monomorphic, lanceolate, oblong to obovate or ovate to oval; stamens exserted, 2–4 mm; filaments pilose basally. |
brown, 2.5–3 mm, glabrous. |
light brown to brown, 2–3 mm, glabrous except for roughened to papillate beak. |
= 40. |
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Flowering year-round. |
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Sandy beaches, dunes, and bluffs or sandy to gravelly inland slopes and flats, coastal grassland and chaparral communities, oak and pine woodlands |
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0-300 m (0-1000 ft) |
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CA
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CO; ID; MT; NE; NV; SD; UT; WY
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The native range of Eriogonum parvifolium is restricted to coastal and near-coastal areas (Los Angeles, Monterey, Orange, San Diego, San Luis Obispo, Santa Barbara, and Ventura counties). The coastal expression (var. parvifolium) has thickened leaf blades (0.5–1.5 × 0.3–0.8 cm) and simple or dichotomous inflorescences of compact clusters of involucres containing white to rose flowers. Highly compact and dense mat-forming plants on rocky bluffs immediately next to the ocean were named var. crassifolium; those with yellow flowers were named var. lucidum. The inland form with thin leaf blades (1.5–3 × 0.3–0.8 cm) and highly-branched, cymose, white-flowered inflorescences is perhaps worthy of continued recognition as var. paynei, although there is no sharp distinction between the extremes. Several expressions of the seacliff wild buckwheat are in cultivation, and unfortunately the California Department of Transportation is using the species in roadside plantings, with the result that it is now established in Santa Clara County. Every effort should be made to halt its introduction beyond its native range. The species is the food plant for two federally endangered butterflies, the El Segundo dotted-blue (Euphilotes battoides allyni), near Los Angeles, and Smith’s dotted-blue (Euphilotes enoptes smithi), near Monterey. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 8 (8 in the flora). Eriogonum brevicaule is highly variable, and the variation has yet to be fully resolved taxonomically. The expressions recognized here will encompass the vast majority of populations. The extreme variation previously under the name var. laxifolium is now reduced with the recognition of var. bannockense (low-elevation or northern phase), var. nanum, and var. caelitum (high-elevation, southern phases). Essentially all of the following species (28–63 below) belong to the Eriogonum brevicaule complex. Eriogonum desertorum, E. loganum, E. spathulatum, E. ostlundii, and E. artificis are allied to the complex associated with var. laxifolium, while E. natum is related to var. cottamii. Eriogonum viridulum and E. ephedroides are allied to E. brevicaule var. brevicaule as are E. contortum and E. acaule. Eriogonum brandegeei is also related, but exactly how is less certain. Allied to this complex of species are on the one hand those related to E. batemanii, and on the other all of the matted perennials belonging to the E. ochrocephalum complex. Essentially all of these species form relatively small populations on discrete edaphic sites and are well isolated one from the other. Unfortunately, a clear separation of E. brevicaule from E. desertorum, E. loganum, and E. spathulatum is not always possible. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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1. Flowering stems and inflorescence branches glabrous | → 2 |
1. Flowering stems and inflorescence branches floccose to tomentose or lanate | → 3 |
2. Plants not pulvinate or cespitose, (0.8-)1.5-5 dm; inflorescences cymose, open, divided 3 times or more; widespread | var. brevicaule |
2. Plants pulvinate and cespitose, 0.3-1.5(-1.8) dm; inflorescences capitate or umbellate to cymose and divided 1-2 times; Utah | var. nanum |
3. Perianths pubescent; se Montana, ne Wyoming | var. canum |
3. Perianths glabrous; sc Idaho, ne Nevada, Utah, sw Wyoming, not se Montana or ne Wyoming | → 4 |
4. Inflorescences divided (2-)3-5 times; perianths usually yellow, rarely ochroleucous | → 5 |
4. Inflorescences capitate or umbellate to cymose and divided 1-2 times; perianths ochroleucous or yellow | → 6 |
5. Flowers (1-)1.5-2.5 mm; sw Wyoming | var. micranthum |
5. Flowers (2.5-)3-4 mm; Utah | var. cottamii |
6. Leaf blades linear to narrowly oblanceolate; inflorescences capitate or divided | → 7 |
6. Leaf blades linear, oblanceolate, or narrowly oblanceolate to elliptic; inflorescences capitate. [8. Shifted to left margin.—Ed.] | → 8 |
7. Leaf blades (1.5-)3-9(-12) × 0.1-0.5(-0.7) cm, tomentose abaxially, less so and grayish or occasionally greenish adaxially, margins usually revolute, occasionally plane; perianths ochroleucous or yellow; n Utah, se Idaho, below 2800 m | var. laxifolium |
7. Leaf blades 0.2-4.5(-5) × (0.2-)0.3-0.6(-0.7) cm, tomentose abaxially, thinly floccose and bright green adaxially, margins plane or slightly thickened; perianths yellow; c Utah, above 2700 m | var. caelitum |
8. Leaf blades and flowering stems bright green under the thinly floccose pubescence; ne Nevada and nw Utah | var. laxifolium |
8. Leaf blades and flowering stems usually grayish to dull greenish under tomentum, rarely thinly floccose; se Idaho and sw Wyoming s to n Utah and ne Nevada | → 9 |
9. Leaf margins crenulate; leaf blades narrowly oblanceolate to narrowly elliptic, (0.3-)0.5-1.5(-2) × 0.2-0.5(-7) cm, densely white-tomentose abaxially, floccose and greenish adaxially | var. nanum |
9. Leaf margins usually plane (rarely crenulate in Wyoming); leaf blades narrowly oblanceolate to oblanceolate, (0.8-)1-4(-4.5) × (0.3-)0.4-0.8 cm, densely tomentose abaxially, tomentose to floccose and grayish to greenish adaxially | var. bannockense |
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FNA vol. 5, p. 300. |
FNA vol. 5, p. 264. |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
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E. parvifolium var. crassifolium, E. parvifolium subsp. lucidum, E. parvifolium var. lucidum, E. parvifolium subsp. paynei, E. parvifolium var. paynei |
E. campanulatum subsp. brevicaule |
Smith: in A. Rees, Cycl. 13(2): Eriogonum no. 2. (1809) |
Nuttall: Proc. Acad. Nat. Sci. Philadelphia 4: 15. (1848) |
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