Flora of North America species comparison

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Eriogonum nudum

Eriogonum heermannii

bare-stem buckwheat, barestem wild buckwheat, naked buckwheat, naked wild buckwheat, nude buckwheat

Heerman buckwheat, Heermann's buckwheat, Heermann's wild buckwheat
Habit Herbs, mostly erect, infrequently scapose, (0.5–)1–15(–20) × 0.5–3 dm, glabrous or floccose to tomentose, usually greenish, occasionally grayish. Shrubs and subshrubs, spreading to rounded and occasionally erect, not scapose, (0.5–)1–20 × 2–25 dm, glabrous or occasionally floccose, sometimes scabrellous, greenish, infrequently grayish.
Stems

spreading to erect, with or without persistent leaf bases, up to 1/4 height of plant;

caudex stems absent;

aerial flowering stems erect to spreading, slender to stout, solid or hollow, occasionally fistulose, 0.3–4(–10) dm, glabrous or lanate to tomentose or floccose.

spreading or erect, without persistent leaf bases, up to 1/2 height of plant;

caudex stems absent;

aerial flowering stems erect or nearly so, slender to stout, solid, not fistulose, 0.02–0.5 dm, thinly tomentose or glabrous.
Leaves

basal or sheathing up stem 0.5–4 dm;

petiole 1–10 cm, glabrous or tomentose;

blade oblanceolate to elliptic or ovate, 1–6 × (0.3–)1–4 cm, densely white-lanate or tomentose abaxially, tomentose to floccose or subglabrous to glabrous adaxially, margins plane or undulate-crisped.

cauline, 1 per node, quickly deciduous;

petiole 0.1–1.5 cm, floccose or glabrous;

blade linear, oblanceolate or spatulate or elliptic, or oblong, (0.4–)1–2(–4) × 0.1–0.8 cm, tomentose to floccose or glabrous abaxially, floccose to thinly floccose or glabrous adaxially, margins plane.
Inflorescences

cymose, rarely umbellate or capitate, 2–100(–150) × 2–40(–80) cm;

branches usually dichotomous, glabrous or tomentose to floccose or sparsely pubescent;

bracts 3, scalelike, 0.5–3(–5) mm.

cymose or racemose, 1–25(–30) × 1–30(–35) cm;

branches dichotomous, sometimes with secondaries suppressed, smooth or angled to ridged and grooved, glabrous or occasionally floccose or scabrous;

bracts 3, scalelike, 0.3–2 mm.
Peduncles

absent.

absent.
Involucres

1 per node or 2–10 per cluster, turbinate to turbinate-campanulate, (2.5–)3–5(–7) × (1.5–)2–4 mm, glabrous, tomentose, or sparsely pubescent;

teeth 5–8, 0.2–0.6 mm.

1 per node, narrowly turbinate or campanulate, 0.7–3 × 0.7–4 mm, glabrous, infrequently floccose;

teeth 5, erect, 0.3–0.7 mm.
Flowers

(1.5–)2–4 mm;

perianth white or yellow, sometimes pink or rose, glabrous or pubescent;

tepals connate proximal 1/4, monomorphic, oblong to obovate;

stamens exserted, 2–5 mm;

filaments pilose proximally.

(1.5–)2–4 mm;

perianth white, yellowish white, pink, or reddish, glabrous;

tepals connate proximal 1/4, dimorphic, those of outer whorl obovate to orbiculate, those of inner whorl narrowly lanceolate to oblong;

stamens exserted, 2–5 mm;

filaments pilose proximally.
Achenes

light brown to brown, 1.5–3.5 mm, glabrous.

light brown to brown, 2–5 mm, glabrous.

Eriogonum nudum

Eriogonum heermannii
Distribution
from FNA
CA; NV; OR; WA; nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NV; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 13 (13 in the flora).

Several local groups of Native Americans in California used members of this species in a variety of ways. S. A. Barrett and E. W. Gifford (1933) and S. M. Schenck and E. W. Gifford (1952) reported the consumption of raw young stems that are rather moist and tasty, although there is a sour aftertaste. The Kawaiisu used the hollow stems (probably var. westonii, rather than var. pauciflorum) as drinking tubes and as pipes (M. L. Zigmond 1981). Zigmond reported also that the roots of var. pauciflorum are used as an infusion for coughs.

Members of Eriogonum nudum are food plants for the Bauer’s dotted-blue butterfly (Euphilotes baueri), the Pacific dotted-blue (E. enoptes), the gorgon copper (Gaeides gorgon), and the Mormon metalmark (Apodemia mormo).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 8 (8 in the flora).

As Eriogonum heermannii is here circumscribed, the number of varieties is dramatically decreased from past presentations, with E. apachense reduced to synonymy under a now greatly expanded var. argense. Also included in that variety is the more stoutly branched var. subracemosum. The southern var. heermannii of basically desert ranges and the more northern var. occidentale of the Coast Ranges in California are maintained, but their separation is more traditional than certain. The fragile and bulky nature of many dried, often poorly prepared specimens, and the tendency for leaves to fall away have made varietal identification within E. heermannii difficult.

Eriogonum heermannii varieties are food plants for Ellis’s dotted-blue butterfly (Euphilotes ellisi) and the Mormon metalmark (Apodemia mormo mormo).

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key
1. Involucres and inflorescence branches tomentose to floccose; leaves usually basal
→ 2
1. Involucres and inflorescence branches glabrous or, if pubescent, leaves sheathing up stems
→ 3
2. Perianths white; flowering stems lanate to tomentose; leaf blades 2-3.5 cm; s Sierra Nevada
var. regirivum
2. Perianths yellow or white to rose; flowering stems tomentose to floccose; leaf blades 2-4 cm; s Oregon, n California, wc Nevada
var. oblongifolium
3. Leaves sheathing up stems, margins often strongly undulate-crisped
→ 4
3. Leaves basal, margins plane or slightly undulate-crisped
→ 8
4. Flowering stems tomentose
→ 5
4. Flowering stems glabrous
→ 6
5. Perianths pubescent; flowers 1.5-2 mm; involucres 3-4 mm; Sierra Nevada, California
var. regirivum
5. Perianths glabrous; flowers 3-4 mm; involucres 4-6 mm; Coast Ranges, California
var. decurrens
6. Leaf blades densely lanate abaxially, tomentose adaxially; involucres 5-10 per cluster; Sierra Nevada, California
var. murinum
6. Leaf blades tomentose abaxially, less so to floccose, glabrous, or nearly so adaxially; involucres 1 per node or 2-5 per cluster; widespread, Coast Ranges, California
→ 7
7. Flowering stems occasionally fistulose; involucres (2-)3-5 per cluster; perianths white to pink, rarely yellowish
var. auriculatum
7. Flowering stems strongly fistulose; involucres 1 per node; perianths pale yellowish white to yellow or white
var. indictum
8. Involucres 1(-2) per cluster
var. westonii
8. Involucres 2-10 per cluster
→ 12
9. Flowering stems fistulose; perianths yellow, infrequently white; c California
var. westonii
9. Flowering stems not fistulose or, if so, plants of sw California; perianths white, rarely yellow
→ 10
10. Perianths pubescent or, infrequently, glabrous; s California
var. pauciflorum
10. Flowers glabrous; sw Oregon, n California, Nevada
→ 11
11. Leaf blades 1-5 cm; sw Oregon, n California (including w slope of Sierra Nevada), Washington
var. nudum
11. Leaf blades 1-2 cm; Sierra Nevada of California and adjacent desert ranges of wc Nevada
var. deductum
12. Perianths pubescent, often yellow
→ 13
12. Perianths usually glabrous abaxially, white, rarely yellow
→ 14
13. Flowering stems not fistulose; leaf blades floccose or glabrous adaxially, margins plane; plants of nonaridregions
var. pubiflorum
13. Flowering stems slightly to distinctly fistulose; leaf blades tomentose to floccose adaxially, margins undulate-crisped; plants of arid regions
var. westonii
14. Inflorescences capitate or nearly so; alpine, Sierra Nevada
E. nudumvar. scapigerum
14. Inflorescences cymose or, if capitate, not alpine.
→ 15
15. Involucres 1(-2) per cluster, 5- 7 mm; mountains of s California
var. pauciflorum
15. Involucres 2-10 per cluster, 3-5 mm; mountains and foothills of c and n California or coastal bluffs. [16. Shifted to left margin—Ed.]
→ 16
16. Inflorescences cymose and branched 2 or more times; involucres 2-5 per cluster; mountains and foothills of c and n California, Oregon, and Washington.
var. nudum
16. Inflorescences capitate or cymose and branched 1-2 times; involucres 5-10 per cluster; coastal bluffs, sw Oregon, ne California
var. paralinum
1. Flowering stems and inflorescence branches round or angled, scabrellous or papillate-scabrous; plants often densely branched
→ 2
1. Flowering stems and inflorescence branches round, not angled, usually smooth, glabrous or thinly tomentose; plants sparsely branched
→ 3
2. Flowering stems and inflorescence branches round, usually distinctly scabrellous or infrequently papillate-scabrous; Arizona, se California, Nevada
var. argense
2. Flowering stems and inflorescence branches sharply ridged and deeply grooved, minutely scabrellous; nw Arizona, se California, s Nevada, sw Utah
var. sulcatum
3. Involucres not racemosely arranged on inflorescence branches, or only last 2-3 so disposed; inflorescences diffusely branched, glabrous; Great Basin or northeast edge of Mojave Desert
→ 4
3. Involucres racemosely arranged along inflorescence branches or at least at tips of branches; inflorescences openly branched, glabrous or floccose to thinly tomentose; Mojave and Sonoran deserts
→ 5
4. Inflorescence branches mostly slender, smooth, not ridged, not spinose; inflorescences 3-15(-23) × 5-20 cm; subshrubs or shrubs, 3-7 × 5-12(-15) dm; Great Basin, ec California and Nevada
var. humilius
4. Inflorescence branches stoutish, faintly ridged and grooved, spinose; inflorescences 3-7(-10) × 3-10(-12) cm; shrubs, 1-3 × 1.5-5(-8) dm; Mojave Desert, sw Utah and nw Arizona
var. subspinosum
5. Inflorescence branches thinly tomentose to floccose; s California, s Nevada, nw Arizona
var. floccosum
5. Inflorescence branches glabrous; s Nevada, California
→ 6
6. Involucres racemosely arranged; inflorescence branches whiplike; s Nevada
var. clokeyi
6. Involucres racemosely arranged only distally; inflorescence branches not whiplike; California
→ 7
7. Leaf blades 0.5-1.5 cm, glabrous abaxially; inflorescence branches stout; sc California
var. heermannii
7. Leaf blades 1.5-3(-4) cm, mostly tomentose to floccose abaxially (at least in early anthesis); inflorescence branches slender; sw California
var. occidentale
Source FNA vol. 5, p. 309. Treatment author: James L. Reveal. FNA vol. 5, p. 304. Treatment author: James L. Reveal.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. nudum var. auriculatum, E. nudum var. decurrens, E. nudum var. deductum, E. nudum var. indictum, E. nudum var. murinum, E. nudum var. nudum, E. nudum var. oblongifolium, E. nudum var. paralinum, E. nudum var. pauciflorum, E. nudum var. pubiflorum, E. nudum var. regirivum, E. nudum var. westonii, E. nudumvar. scapigerum
E. heermannii var. argense, E. heermannii var. clokeyi, E. heermannii var. floccosum, E. heermannii var. heermannii, E. heermannii var. humilius, E. heermannii var. occidentale, E. heermannii var. subspinosum, E. heermannii var. sulcatum
Synonyms E. latifolium subsp. nudum E. geniculatum
Name authority Douglas ex Bentham: Trans. Linn. Soc. London 17: 413. (1836) Durand & Hilgard: in War Department [U.S.], Pacif. Railr. Rep. 5(3): 14. (1857)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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