Flora of North America species comparison

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Eriogonum nudum

Eriogonum exaltatum

bare-stem buckwheat, barestem wild buckwheat, naked buckwheat, naked wild buckwheat, nude buckwheat

ladder buckwheat, ladder wild buckwheat
Habit Herbs, mostly erect, infrequently scapose, (0.5–)1–15(–20) × 0.5–3 dm, glabrous or floccose to tomentose, usually greenish, occasionally grayish. Herbs, erect, annual, (0.5–)3–6(–10) dm, glabrous, glaucous, grayish.
Stems

spreading to erect, with or without persistent leaf bases, up to 1/4 height of plant;

caudex stems absent;

aerial flowering stems erect to spreading, slender to stout, solid or hollow, occasionally fistulose, 0.3–4(–10) dm, glabrous or lanate to tomentose or floccose.

caudex absent;

aerial flowering stems erect, solid, not fistulose, (0.5–)1–2 dm, glabrous.
Leaves

basal or sheathing up stem 0.5–4 dm;

petiole 1–10 cm, glabrous or tomentose;

blade oblanceolate to elliptic or ovate, 1–6 × (0.3–)1–4 cm, densely white-lanate or tomentose abaxially, tomentose to floccose or subglabrous to glabrous adaxially, margins plane or undulate-crisped.

basal;

petiole 1–10 cm, tomentose;

blade sub-cordate to orbiculate, (1.5–)2–5(–8) × (1.5–)2–5(–8) cm, densely white-tomentose abaxially, floccose to subglabrous and green adaxially, margins often wavy.
Inflorescences

cymose, rarely umbellate or capitate, 2–100(–150) × 2–40(–80) cm;

branches usually dichotomous, glabrous or tomentose to floccose or sparsely pubescent;

bracts 3, scalelike, 0.5–3(–5) mm.

cymose, spreading when immature, quickly becoming narrowly erect and strict with whiplike branches, (5–)10–50(–80) × 10–50 cm;

branches glabrous, glaucous;

bracts 3, scalelike, 1–1.5(–2) × 0.4–1 mm.
Peduncles

absent.

absent or erect, straight, slender, 0.1–0.2 cm, glabrous.
Involucres

1 per node or 2–10 per cluster, turbinate to turbinate-campanulate, (2.5–)3–5(–7) × (1.5–)2–4 mm, glabrous, tomentose, or sparsely pubescent;

teeth 5–8, 0.2–0.6 mm.

turbinate, 2–2.5(–3) × 1.5–2.5 mm, glabrous;

teeth 5, erect, 0.5–1.5 mm.
Flowers

(1.5–)2–4 mm;

perianth white or yellow, sometimes pink or rose, glabrous or pubescent;

tepals connate proximal 1/4, monomorphic, oblong to obovate;

stamens exserted, 2–5 mm;

filaments pilose proximally.

1.5–2 mm;

perianth white with green or reddish midribs, becoming pinkish, glabrous;

tepals dimorphic, those of outer whorl oblong, those of inner whorl lanceolate;

stamens exserted, 1.5–2.5 mm;

filaments pilose proximally.
Achenes

light brown to brown, 1.5–3.5 mm, glabrous.

dark brown to blackish, 3-gonous, 2–2.5 mm, glabrous.
2n

= 40.

Eriogonum nudum

Eriogonum exaltatum
Phenology Flowering May–Oct.
Habitat Sandy to gravelly flats and slopes, saltbush, creosote bush, greasewood, blackbrush, and mesquite communities, rarely in pinyon-juniper woodlands
Elevation 500-1400 m (1600-4600 ft)
Distribution
from FNA
CA; NV; OR; WA; nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NV; UT
[BONAP county map]
Discussion

Varieties 13 (13 in the flora).

Several local groups of Native Americans in California used members of this species in a variety of ways. S. A. Barrett and E. W. Gifford (1933) and S. M. Schenck and E. W. Gifford (1952) reported the consumption of raw young stems that are rather moist and tasty, although there is a sour aftertaste. The Kawaiisu used the hollow stems (probably var. westonii, rather than var. pauciflorum) as drinking tubes and as pipes (M. L. Zigmond 1981). Zigmond reported also that the roots of var. pauciflorum are used as an infusion for coughs.

Members of Eriogonum nudum are food plants for the Bauer’s dotted-blue butterfly (Euphilotes baueri), the Pacific dotted-blue (E. enoptes), the gorgon copper (Gaeides gorgon), and the Mormon metalmark (Apodemia mormo).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eriogonum insigne has been a troublesome taxon.  The type, collected by E. Palmer in 1876, is a curious specimen from near Paragonah in Iron County, Utah.  That expression has not been recollected and, after another frustrating review of the type material, it is now referred to E. deflexum, along with other odd specimens from southern Nevada and California that have always been difficult to assign.  As a result, the tall, upright plants with long, whiplike branches of northwestern Arizona (Mohave County), southern Nevada (northeastern Clark and southern Lincoln counties), and southwestern Utah (southwestern Washington County), are now recognized under the name E. exaltatum.  Some specimens remain problematic.  Immature plants of E. exaltatum (Goodding 2302, GH, MIN, MO, NY, RM, UC) resemble E. bifurcatum. Specimens from Eureka Valley (DeDecker 4741, NY) and western Pahrump Valley (Abrams 14248, DS, GH, NY) in Inyo County, California, appear to belong to E. exaltatum.

Like Eriogonum exaltatum, some populations of E. deflexum var. deflexum have nearly erect involucres.  The plants from the Buried Hills (Nye and Clark counties, Nevada) are particularly odd.  Single plants from a few collections in Kane County, Utah, have sessile, seemingly erect involucres in the forks of inflorescence branches.  None of these plants has the characteristic long, whiplike branches of E. exaltatum, and they are not included in E. deflexum var. deflexum. The type of E. deflexum var. rectum from San Bernardino County, California (Reveal & Broome 6385, CAS, NY, US, etc.), has individual specimens with both deflexed and somewhat erect involucres, along with whiplike branches.  Collections from Imperial and San Diego counties, previously associated with what is here termed E. exaltatum, are now considered specimens of E. deflexum.  Clearly, more work is required to understand these species fully.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key
1. Involucres and inflorescence branches tomentose to floccose; leaves usually basal
→ 2
1. Involucres and inflorescence branches glabrous or, if pubescent, leaves sheathing up stems
→ 3
2. Perianths white; flowering stems lanate to tomentose; leaf blades 2-3.5 cm; s Sierra Nevada
var. regirivum
2. Perianths yellow or white to rose; flowering stems tomentose to floccose; leaf blades 2-4 cm; s Oregon, n California, wc Nevada
var. oblongifolium
3. Leaves sheathing up stems, margins often strongly undulate-crisped
→ 4
3. Leaves basal, margins plane or slightly undulate-crisped
→ 8
4. Flowering stems tomentose
→ 5
4. Flowering stems glabrous
→ 6
5. Perianths pubescent; flowers 1.5-2 mm; involucres 3-4 mm; Sierra Nevada, California
var. regirivum
5. Perianths glabrous; flowers 3-4 mm; involucres 4-6 mm; Coast Ranges, California
var. decurrens
6. Leaf blades densely lanate abaxially, tomentose adaxially; involucres 5-10 per cluster; Sierra Nevada, California
var. murinum
6. Leaf blades tomentose abaxially, less so to floccose, glabrous, or nearly so adaxially; involucres 1 per node or 2-5 per cluster; widespread, Coast Ranges, California
→ 7
7. Flowering stems occasionally fistulose; involucres (2-)3-5 per cluster; perianths white to pink, rarely yellowish
var. auriculatum
7. Flowering stems strongly fistulose; involucres 1 per node; perianths pale yellowish white to yellow or white
var. indictum
8. Involucres 1(-2) per cluster
var. westonii
8. Involucres 2-10 per cluster
→ 12
9. Flowering stems fistulose; perianths yellow, infrequently white; c California
var. westonii
9. Flowering stems not fistulose or, if so, plants of sw California; perianths white, rarely yellow
→ 10
10. Perianths pubescent or, infrequently, glabrous; s California
var. pauciflorum
10. Flowers glabrous; sw Oregon, n California, Nevada
→ 11
11. Leaf blades 1-5 cm; sw Oregon, n California (including w slope of Sierra Nevada), Washington
var. nudum
11. Leaf blades 1-2 cm; Sierra Nevada of California and adjacent desert ranges of wc Nevada
var. deductum
12. Perianths pubescent, often yellow
→ 13
12. Perianths usually glabrous abaxially, white, rarely yellow
→ 14
13. Flowering stems not fistulose; leaf blades floccose or glabrous adaxially, margins plane; plants of nonaridregions
var. pubiflorum
13. Flowering stems slightly to distinctly fistulose; leaf blades tomentose to floccose adaxially, margins undulate-crisped; plants of arid regions
var. westonii
14. Inflorescences capitate or nearly so; alpine, Sierra Nevada
E. nudumvar. scapigerum
14. Inflorescences cymose or, if capitate, not alpine.
→ 15
15. Involucres 1(-2) per cluster, 5- 7 mm; mountains of s California
var. pauciflorum
15. Involucres 2-10 per cluster, 3-5 mm; mountains and foothills of c and n California or coastal bluffs. [16. Shifted to left margin—Ed.]
→ 16
16. Inflorescences cymose and branched 2 or more times; involucres 2-5 per cluster; mountains and foothills of c and n California, Oregon, and Washington.
var. nudum
16. Inflorescences capitate or cymose and branched 1-2 times; involucres 5-10 per cluster; coastal bluffs, sw Oregon, ne California
var. paralinum
Source FNA vol. 5, p. 309. FNA vol. 5, p. 397.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. nudum var. auriculatum, E. nudum var. decurrens, E. nudum var. deductum, E. nudum var. indictum, E. nudum var. murinum, E. nudum var. nudum, E. nudum var. oblongifolium, E. nudum var. paralinum, E. nudum var. pauciflorum, E. nudum var. pubiflorum, E. nudum var. regirivum, E. nudum var. westonii, E. nudumvar. scapigerum
Synonyms E. latifolium subsp. nudum
Name authority Douglas ex Bentham: Trans. Linn. Soc. London 17: 413. (1836) M. E. Jones: Contr. W. Bot. 15: 61. (1929)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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