Flora of North America species comparison

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Eriogonum nudum

Eriogonum corymbosum

bare-stem buckwheat, barestem wild buckwheat, naked buckwheat, naked wild buckwheat, nude buckwheat

corymbed wild buckwheat, crispleaf buckwheat, crispleaf wild buckwheat, white wild buckwheat
Habit Herbs, mostly erect, infrequently scapose, (0.5–)1–15(–20) × 0.5–3 dm, glabrous or floccose to tomentose, usually greenish, occasionally grayish. Shrubs or subshrubs, spreading, rounded, occasionally erect, rarely somewhat matted, not scapose, (0.5–)1.5–8(–15) × (2–)3–15(–23) dm, grayish- to reddish-brown-tomentose to floccose or glabrous, grayish or greenish.
Stems

spreading to erect, with or without persistent leaf bases, up to 1/4 height of plant;

caudex stems absent;

aerial flowering stems erect to spreading, slender to stout, solid or hollow, occasionally fistulose, 0.3–4(–10) dm, glabrous or lanate to tomentose or floccose.

spreading or erect, often with persistent leaf bases, up to 3/4 or more height of plant;

caudex stems absent or somewhat matted;

aerial flowering stems erect or nearly so, slender or occasionally stout, solid, not fistulose, (0.1–)1–2 dm, tomentose to floccose, occasionally glabrous.
Leaves

basal or sheathing up stem 0.5–4 dm;

petiole 1–10 cm, glabrous or tomentose;

blade oblanceolate to elliptic or ovate, 1–6 × (0.3–)1–4 cm, densely white-lanate or tomentose abaxially, tomentose to floccose or subglabrous to glabrous adaxially, margins plane or undulate-crisped.

cauline, 1 per node;

petiole 0.1–1.5 cm, tomentose to floccose;

blade lanceolate to oblanceolate or elliptic to nearly orbiculate, rarely cordate, (0.5–)1–3(–4.5) × (0.3–)0.5–3(–3.5) cm, densely white-, tannish- or brownish-tomentose on both surfaces or less so to nearly glabrous and green adaxially, margins occasionally crenulate.
Inflorescences

cymose, rarely umbellate or capitate, 2–100(–150) × 2–40(–80) cm;

branches usually dichotomous, glabrous or tomentose to floccose or sparsely pubescent;

bracts 3, scalelike, 0.5–3(–5) mm.

cymose, rarely capitate or umbellate, diffuse to rather open, (1–)3–20 × 2–25(–30) cm;

branches dichotomous, tomentose, floccose, or rarely glabrous;

bracts 3, scalelike, usually triangular, and 1–3(–6) mm, or leaflike, 10–25 mm, and similar to leaf blades.
Peduncles

absent.

absent.
Involucres

1 per node or 2–10 per cluster, turbinate to turbinate-campanulate, (2.5–)3–5(–7) × (1.5–)2–4 mm, glabrous, tomentose, or sparsely pubescent;

teeth 5–8, 0.2–0.6 mm.

1 per node, turbinate, (1–)1.5–3.5 × 1–2(–2.5) mm;

teeth 5, erect, 0.3–1 mm.
Flowers

(1.5–)2–4 mm;

perianth white or yellow, sometimes pink or rose, glabrous or pubescent;

tepals connate proximal 1/4, monomorphic, oblong to obovate;

stamens exserted, 2–5 mm;

filaments pilose proximally.

(1.5–)2–3.5 mm;

perianth white to cream, pink, or pale yellow to yellow, glabrous or rarely sparsely pilose;

tepals connate proximal 1/4–1/3, essentially monomorphic, oblanceolate to spatulate;

stamens included to slightly exserted, 1–4(–5) mm;

filaments typically pilose proximally.
Achenes

light brown to brown, 1.5–3.5 mm, glabrous.

brown, 2–2.5(–3) mm, glabrous except for occasional papillate beak.

Eriogonum nudum

Eriogonum corymbosum
Distribution
from FNA
CA; NV; OR; WA; nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; NM; NV; TX; UT; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 13 (13 in the flora).

Several local groups of Native Americans in California used members of this species in a variety of ways. S. A. Barrett and E. W. Gifford (1933) and S. M. Schenck and E. W. Gifford (1952) reported the consumption of raw young stems that are rather moist and tasty, although there is a sour aftertaste. The Kawaiisu used the hollow stems (probably var. westonii, rather than var. pauciflorum) as drinking tubes and as pipes (M. L. Zigmond 1981). Zigmond reported also that the roots of var. pauciflorum are used as an infusion for coughs.

Members of Eriogonum nudum are food plants for the Bauer’s dotted-blue butterfly (Euphilotes baueri), the Pacific dotted-blue (E. enoptes), the gorgon copper (Gaeides gorgon), and the Mormon metalmark (Apodemia mormo).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 8 (8 in the flora).

Eriogonum corymbosum is a difficult complex of overlapping expressions, some of which are maintained here as taxonomically significant. Although perianth color is used to group the varieties, this feature is not consistent even in single populations. Therefore, population trends in perianth color must be noted in the field. Most of the varieties are then distinguished on the basis of leaf characters, and again, considerable variation can be seen in some populations. Still, the combination of flower color, leaf features, and geographic distribution should prove useful in distinguishing the varieties.

S. L. Welsh et al. (2003) alluded to hybrid combinations involving Eriogonum corymbosum and other species. Aside from the instances involving E. brevicaule, discussed below, none has been confirmed. Most of the putative hybrids are misidentified specimens of E. lonchophyllum or collections of var. corymbosum in which the leaf-margins are not decidedly crisped, a feature usually seen only in fully mature plants.

Eriogonum corymbosum was widely used by Native Americans. P. A. Vestal (1940) reported that the Hopi pressed boiled stalks into cakes that, when dried, were eaten with salt. J. W. Fewkes (1896) indicated that boiled leaves were mixed with cornmeal and water, and then baked into a kind of bread. S. A. Weber and P. D. Seaman (1985) indicated that A. F. Whiting was aware of a decoction of leaves (probably from var. glutinosum) being used for headaches. Variety glutinosum also was used primarily to treat tuberculosis, or at least as a cough medicine (D. E. Moerman 1986).

Some of the expressions of Eriogonum corymbosum are attracting the interest of gardeners, a few are coming into cultivation, and several selections are now being developed. The plants are slow growing but can be transplanted with some degree of success. Members of the varieties are food plants for Ellis’s dotted-blue butterfly (Euphilotes ellisi).

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key
1. Involucres and inflorescence branches tomentose to floccose; leaves usually basal
→ 2
1. Involucres and inflorescence branches glabrous or, if pubescent, leaves sheathing up stems
→ 3
2. Perianths white; flowering stems lanate to tomentose; leaf blades 2-3.5 cm; s Sierra Nevada
var. regirivum
2. Perianths yellow or white to rose; flowering stems tomentose to floccose; leaf blades 2-4 cm; s Oregon, n California, wc Nevada
var. oblongifolium
3. Leaves sheathing up stems, margins often strongly undulate-crisped
→ 4
3. Leaves basal, margins plane or slightly undulate-crisped
→ 8
4. Flowering stems tomentose
→ 5
4. Flowering stems glabrous
→ 6
5. Perianths pubescent; flowers 1.5-2 mm; involucres 3-4 mm; Sierra Nevada, California
var. regirivum
5. Perianths glabrous; flowers 3-4 mm; involucres 4-6 mm; Coast Ranges, California
var. decurrens
6. Leaf blades densely lanate abaxially, tomentose adaxially; involucres 5-10 per cluster; Sierra Nevada, California
var. murinum
6. Leaf blades tomentose abaxially, less so to floccose, glabrous, or nearly so adaxially; involucres 1 per node or 2-5 per cluster; widespread, Coast Ranges, California
→ 7
7. Flowering stems occasionally fistulose; involucres (2-)3-5 per cluster; perianths white to pink, rarely yellowish
var. auriculatum
7. Flowering stems strongly fistulose; involucres 1 per node; perianths pale yellowish white to yellow or white
var. indictum
8. Involucres 1(-2) per cluster
var. westonii
8. Involucres 2-10 per cluster
→ 12
9. Flowering stems fistulose; perianths yellow, infrequently white; c California
var. westonii
9. Flowering stems not fistulose or, if so, plants of sw California; perianths white, rarely yellow
→ 10
10. Perianths pubescent or, infrequently, glabrous; s California
var. pauciflorum
10. Flowers glabrous; sw Oregon, n California, Nevada
→ 11
11. Leaf blades 1-5 cm; sw Oregon, n California (including w slope of Sierra Nevada), Washington
var. nudum
11. Leaf blades 1-2 cm; Sierra Nevada of California and adjacent desert ranges of wc Nevada
var. deductum
12. Perianths pubescent, often yellow
→ 13
12. Perianths usually glabrous abaxially, white, rarely yellow
→ 14
13. Flowering stems not fistulose; leaf blades floccose or glabrous adaxially, margins plane; plants of nonaridregions
var. pubiflorum
13. Flowering stems slightly to distinctly fistulose; leaf blades tomentose to floccose adaxially, margins undulate-crisped; plants of arid regions
var. westonii
14. Inflorescences capitate or nearly so; alpine, Sierra Nevada
E. nudumvar. scapigerum
14. Inflorescences cymose or, if capitate, not alpine.
→ 15
15. Involucres 1(-2) per cluster, 5- 7 mm; mountains of s California
var. pauciflorum
15. Involucres 2-10 per cluster, 3-5 mm; mountains and foothills of c and n California or coastal bluffs. [16. Shifted to left margin—Ed.]
→ 16
16. Inflorescences cymose and branched 2 or more times; involucres 2-5 per cluster; mountains and foothills of c and n California, Oregon, and Washington.
var. nudum
16. Inflorescences capitate or cymose and branched 1-2 times; involucres 5-10 per cluster; coastal bluffs, sw Oregon, ne California
var. paralinum
1. Perianths usually pale yellow to yellow, occasionally white; s Utah, n Arizona, se Nevada
→ 2
1. Perianths white to cream or pink; sw Wyoming s through e Utah and w Colorado to nw New Mexico and Arizona
→ 4
2. Inflorescence branches glabrous or nearly so; sw Utah
var. aureum
2. Inflorescence branches tomentose to floccose
→ 3
3. Leaf blades greenish-floccose adaxially; Colorado Plateau, n Arizona, s Utah
var. glutinosum
3. Leaf blades silvery-floccose adaxially; Mojave Desert, se Nevada, sc Utah
var. nilesii
4. Leaf blades elliptic-oblong to nearly orbiculate, 1-3(-3.5) × 1-2.5(-3.5) cm
→ 5
4. Leaf blades lanceolate to oblanceolate or elliptic, (0.5-)1-3(-4.5) × (0.3-)0.5-1.5 cm
→ 6
5. Leaf blades elliptic-oblong to oblong or ovate, rarely cordate, densely white- to tan-lanate or less so to brownish-floccose adaxially; plants brownish-white- tomentose; flowers 2-2.5(-3) mm
var. velutinum
5. Leaf blades usually broadly ovate to nearly orbiculate, densely grayish-tomentose abaxially, sometimes tomentose to floccose on both surfaces, or more often subglabrous or glabrous and green adaxially; plants greenish-tomentose; flowers 2.5-3 mm
var. orbiculatum
6. Plants sprawling subshrubs, 0.5-1.5 dm, occasionally shrubs, 2-4.5 dm; leaf blades 0.5-2 × 0.3-0.6(-0.8) cm; subalpine in bristlecone pine communities
var. heilii
6. Plants erect subshrubs or shrubs, 3-10 dm; leaf blades 1-3(-4.5) × (0.3-)0.5-1.5 cm; desert scrub to montane forest communities
→ 7
7. Leaves cauline 2 or more length of flowering stems; n Arizona, w Colorado, e and s Utah, sw Wyoming
var. corymbosum
7. Leaves cauline 4 or less length of flowering stems; c and e Utah
var. revealianum
Source FNA vol. 5, p. 309. FNA vol. 5, p. 255.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. nudum var. auriculatum, E. nudum var. decurrens, E. nudum var. deductum, E. nudum var. indictum, E. nudum var. murinum, E. nudum var. nudum, E. nudum var. oblongifolium, E. nudum var. paralinum, E. nudum var. pauciflorum, E. nudum var. pubiflorum, E. nudum var. regirivum, E. nudum var. westonii, E. nudumvar. scapigerum
E. corymbosum var. aureum, E. corymbosum var. corymbosum, E. corymbosum var. glutinosum, E. corymbosum var. heilii, E. corymbosum var. nilesii, E. corymbosum var. orbiculatum, E. corymbosum var. revealianum, E. corymbosum var. velutinum
Synonyms E. latifolium subsp. nudum E. effusum subsp. corymbosum
Name authority Douglas ex Bentham: Trans. Linn. Soc. London 17: 413. (1836) Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 14: 17. (1856)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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