slender buckwheat, slender wild buckwheat
|
wild buckwheat
|
Subshrubs or shrubs, erect to spreading, not scapose, 0.2–1.5 × (0.6–)1–13(–16) dm, white- to tannish-tomentose, floccose, or glabrous. |
Shrubs, subshrubs, or herbs, sometimes nearly arborescent (Eriogonum), perennial, biennial, or annual, homophyllous, polycarpic (rarely monocarpic in Eriogonum); taproot solid or, rarely, chambered (Eriogonum), slender to stout. |
spreading to erect, typically without persistent leaf bases, up to 1/2 height of plant; caudex stems absent or spreading; aerial flowering stems erect to spreading, slender, solid, not fistulose, 0.05–1.5 dm, lanate, tomentose, floccose, subglabrous, or glabrous. |
prostrate or decumbent to spreading or erect, sometimes scapose, rarely absent (Eriogonum), without recurved spines, glabrous or pubescent, sometimes glandular; nodes not swollen; tendrils absent; caudex stems tightly compact to spreading and at or just below the soil surface or spreading to erect and above the soil surface, woody; aerial flowering stems decumbent to spreading or erect, arising at nodes of caudex branches, at distal nodes of aerial branches, or directly from the root, slender to stout and solid or slightly to distinctly fistulose, rarely disarticulating in ringlike segments (Eriogonum). |
cauline, 1 per node or fasciculate; petiole 0.1–0.5 cm, tomentose to floccose or glabrous; blade usually elliptic, sometimes linear to obovate, 0.3–3.5 × (0.07–)0.1–1.2 cm, tomentose abaxially, less so or glabrous adaxially, margins occasionally revolute. |
deciduous (persistent in some shrubby and matted Eriogonum species), basal or basal and cauline, rarely only cauline, rosulate, alternate, or infrequently opposite (Goodmania) or in whorls of 3 (Gilmania); stipules absent (possibly vestigial in some perennial species of Chorizanthe); petiole present, sometimes indistinct, not articulate or with extrafloral nectaries; blade simple, rarely lobed (Pterostegia), rarely awn-tipped (Goodmania). |
cymose, compact, often flat-topped, 0.5–6(–12) × 1–10(–13) cm; branches dichotomous, whitish-lanate to brownish- or reddish-tomentose to floccose or glabrate, infrequently green or gray and subglabrous or glabrous; bracts 3, scalelike, linear to triangular, 1–5 mm. |
terminal or terminal and axillary, cymose and dichotomously or trichotomously branched, or racemose, simple or compound umbellate, or capitate; bracts usually connate proximally, leaflike or scalelike, entire apically, sometimes awn-tipped, glabrous or pubescent. |
absent or mostly erect, slender, 0.3–1.5 cm, tomentose to floccose. |
absent or erect to deflexed relative to inflorescence branch, sometimes reflexed, straight or curved. |
1 per node, turbinate, (1.5–)2–3.5(–4) × 1.3–2.5(–3) mm, tomentose, floccose, subglabrous, or glabrous; teeth 5, erect, (0.3–)0.5–1(–1.7) mm. |
|
1.5–3(–4) mm; perianth yellow or white to pink, orange, rose, red, or occasionally cream, glabrous; tepals connate proximal 1/5–2/5, essentially monomorphic, oblong to obovate; stamens usually exserted, 2.5–4 mm; filaments sparsely to densely puberulent proximally. |
(1–)2–30(–100) per involucral structure, occasionally with stipelike base distal to articulations (Eriogonum); perianth accrescent in fruit, mostly white to red, yellow, light green, greenish white, maroon, or purple, urceolate to campanulate, occasionally glandular or pustulose abaxially, nearly always minutely glandular along midvein adaxially, glabrous or pubescent; tepals (5–)6, in 2 whorls of 3, connate proximally, typically not forming tube (except Chorizanthe, Lastarriaea, Mucronea, Pterostegia), petaloid or, rarely, coriaceous (Lastarriaea), monomorphic or dimorphic, entire, emarginate, or lobed to laciniate apically, rarely awn-tipped (Lastarriaea) or apiculate (Eriogonum); nectary a disk at base of ovary; stamens 3, 6, or 9 (variously 3–9 in Chorizanthe, Mucronea); staminodes absent; filaments usually distinct, occasionally forming staminal tube (Chorizanthe); pistils 3-carpellate, homostylous; ovary 1-locular; ovule 1, orthotropous, placentation basal; styles 3, distinct; stigmas capitate. |
brown, 1.5–3 mm, glabrous. |
brown to black or maroon, homocarpic, winged or unwinged, 3-gonous, less often lenticular or globose-lenticular to globose. |
|
embryo straight or curved. |
|
structures tubular (involucre) or consisting of a series of individual bractlike lobes (involucral bracts) arranged in whorls or spirals, rarely absent (Gilmania), awns present or absent; involucre cylindric, prismatic, turbinate, campanulate, urceolate, or funnelform with 3–8(–36) usually erect teeth or 4–12 spreading to reflexed lobes (teeth and lobes are distal portions of proximally connate involucral bracts); involucral bracts in 1–3 whorls, rarely in spirals (Johanneshowellia), free or connate only at base, linear to oblanceolate or ovate. |
|
|
|
|
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY
|
Mainly temperate regions of w North America (Alaska to Mexico); uncommon in South America (Argentina and Chile) and e North America (WVa s to c Fla, e to Mo, Okla, and Tex) |
Varieties 13 (13 in the flora). Eriogonum microthecum is used as browse by deer and to a lesser degree by cattle and sheep. Some forms are now in cultivation. The species is reportedly used by the Piute of Nevada in the treatment of tuberculosis, lameness, rheumatism, and bladder trouble (P. Train et al. 1941). S. A. Weber and P. D. Seaman (1985) stated that A. F. Whiting found the plants being used as a tea by the Havasupai in northern Arizona. Members of E. microthecum are food plants for subspecies of the rare pallid blue butterfly (Euphilotes pallescens). Also found on this species is the cythera metalmark (Apodemia mormo cythera). Some authors have referred E. effusum to this species, even though the ranges of the two species do not overlap and intermediates are unknown. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 20, species ca. 325 (19 genera, 281 species in the flora). Detailed habitat, elevation, and distribution data for the eriogonoid genera are maintained by the author and available on the Web at: “Eriogonoideae (Polygonaceae) of North America north of Mexico” (http://www.life.umd.edu/emeritus/reveal/pbio/eriog/key.html). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| → 2 |
1. Perianths various shades of white, cream, orange, pink, or red, not yellow | → 4 |
2. Flowering stems and inflorescence branches usually glabrous; e Oregon and wc Idaho | var. microthecum |
2. Flowering stems and inflorescence branches tomentose to floccose; se Oregon and sw Idaho s to e California and w Nevada | → 3 |
3. Leaf blades (0.2-)0.3-0.6(-0.8) cm wide; flowers (1.5-)2-2.5(-3) mm; involucres 2-2.5 mm; achenes 1.5-2 mm; se Oregon and sw Idaho s to e California and w Nevada | var. ambiguum |
3. Leaf blades 0.5-1.2 cm wide; flowers 2.5-3 mm; involucres 2.5-4 mm; achenes 2.5-3 mm; ne California, nw Nevada | var. schoolcraftii |
4. Tomentum whitish (see also var. alpinum of the Sierra Nevada, California); flowering stems and inflorescence branches infrequently glabrous | → 5 |
4. Tomentum brownish or reddish (may be white in var. alpinum), or flowering stems and inflorescence branches essentially glabrous | → 6 |
5. Leaf margins not revolute; flowering inflorescence branches floccose or glabrous; northern phase of species | var. laxiflorum |
5. Leaf margins revolute or nearly so; flowering inflorescence branches lanate to tomentose, or if subglabrous or glabrous, then southern phase of species | var. simpsonii |
| → 7 |
6. Plants subshrubs, 0.2-1.5(-2) dm | → 8 |
7. Flowering stems and inflorescence branches tomentose when young, becoming floccose at maturity; flowers 1.5-2(-2.5) mm; achenes 1.8-2 mm; Death Valley region, California | var. panamintense |
7. Flowering stems and inflorescence branches lanate to tomentose at maturity; flowers 2-2.5(-3) mm; achenes 2.5-3 mm; Transverse Ranges, California | E. microthecumvar. corymbosoides |
8. Leaf blades elliptic or ovate, margins not revolute; flowers (1.5-)2-3.5(-4) mm | → 9 |
8. Leaf blades linear or linear-oblanceolate to narrowly elliptic, margins often revolute; flowers 1.5-2.5 mm | → 10 |
9. Leaf blades 0.5-1 × (0.2-)0.3-0.5(-0.6) cm; involucres (2-)2.5-3 mm; flowers (2.5-)3-3.5(-4) mm; San Gabriel Mountains, California | var. johnstonii |
9. Leaf blades 0.3-0.7(-0.8) × 0.1-0.4 cm; involucres 2.5-3.5 mm; flowers (1.5-)2-3.5 mm; desert ranges of se California, c Nevada, and Utah1f. Eriogonum microthecum var. lapidicola [8. Shifted to left margin—Ed.] | → 8 |
10. Involucres 3-4 mm; perianths cream; San Bernardino Mountains, California | var. lacus-ursi |
10. Involucres (1.5-)2-3 mm; perianths white, pink, red, or rose; Sierra Nevada of California, or e Nevada and w Utah | → 11 |
11. Flowering stems glabrous; leaf blades sparsely floccose or glabrous adaxially; desert ranges, se Nevada | var. arceuthinum |
11. Flowering stems white- to brownish-floccose to subglabrous, or reddish-tomentose to floccose; leaf blades floccose to subglabrous adaxially; California or wc Utah | → 12 |
12. Tomentum white to brownish; Sierra Nevada, California | var. alpinum |
12. Tomentum reddish; desert ranges, wc Utah | var. phoeniceum |
|
1. Plants annual with involucre forming distinct tube, this cylindric or prismatic and awn-tipped typically with at least some awns uncinate or curved, rarely turbinate or urceolate to campanulate and then with 5 divergent, straight awns, or reduced to 1-4, often awn-tipped involucral bracts; flowers mostly 1-2(-6) per involucre | → 2 |
1. Plants perennial or biennial, or, if annual, with involucre forming distinct tube, this turbinate to campanulate or hemispheric and awnless or tipped with erect, straight awns or reduced to awnless involucral bracts, absent entirely in Gilmania; flowers mostly (2-)6-200 per involucre (Eriogoneae, Eriogonineae) | → 10 |
2. Involucres reduced to 1 highly modified, 2-winged bract; leaf blades entire or variously lobed; stems sprawling and spreading (Pterostegieae) | Pterostegia |
2. Involucres tubular or reduced to 3(-4) bracts; leaf blades entire; stems erect to prostrate | → 3 |
3. Involucres reduced to 3(-4) bracts; tepals mucronate or awn-tipped; California | → 4 |
3. Involucres tubular; tepals not mucronate or awn-tipped (Chorizanthineae); widespread | → 5 |
4. Perianths yellowish, densely tomentose abaxially; tepals mucronate apically; stamens 6 or 9; inflorescence branches tomentose (Hollisteriineae) | Hollisteria |
4. Perianths light green to greenish white, thinly pubescent abaxially; tepals acute or awn-tipped apically; stamens 3; inflorescence branches thinly pubescent (Chorizanthineae) | Lastarriaea |
5. Inflorescence bracts opposite, mostly 2, sometimes numerous and whorled; flowers 1(-2) per involucre; w North America | Chorizanthe |
5. Inflorescence bracts alternate and positioned on one side of branch or perfoliate around branch, 3-lobed or parted; flowers (1-)2-6 per involucre; widespread | → 6 |
6. Involucres 5-toothed, each terminated by divergent awn; flowers 4(-6) per involucre; perianth pubescent abaxially; wc California | Aristocapsa |
6. Involucres 2-4(6)-lobed or -toothed, or, if 5-awned apically, involucre with additional basal awns; flowers (1-)2-3 per involucre, perianth glabrous by densely papillate or pubescent abaxially; California | → 7 |
7. Involucres not awned basally; California | → 8 |
7. Involucres awned basally; sw North America | → 9 |
8. Perianths pubescent abaxially; flowers bisexual | Mucronea |
8. Perianths glabrous, densely papillate abaxially; flowers bisexual and unisexual, with proximal 1 pistillate and distal 1 bisexual | Systenotheca |
9. Basal awns 3, on saccate lobes; terminal awns 5, straight, involucres 3-angled; flowers 2 per involucre; sw North America | Centrostegia |
9. Basal awns 6, on nonsaccate basal lobes; terminal awns 6, uncinate; involucres 6-angled; flowers 3 per involucre; sw California | Dodecahema |
10. Plants perennial, if annual or biennial involucre tubular and awnless | → 11 |
10. Plants annual; involucres tubular and awn-tipped or series of free or basally connate, awnless bracts, rarely absent (Gilmania) | → 12 |
11. Involucres tubular, the lobes rarely connate proximally; plants annual, biennial, or perennial; widespread | Eriogonum |
11. Involucres reduced to series of 2-5 obscure, awnless bracts; plants perennial shrubs; ec California | Dedeckera |
12. Involucres not awned or absent | → 13 |
12. Involucres awn-tipped | → 16 |
13. Involucral bracts in 2 whorls of 3; sw Wyoming to ne Arizona and nw New Mexico | Stenogonum |
13. Involucral bracts absent or in 1 whorl or tight spiral; California, Nevada, sw Utah | → 14 |
14. Stamens 3; s California, sw Arizona | Nemacaulis |
14. Stamens 9; se California, s Nevada, sw Utah | → 15 |
15. Perianths thinly pubescent abaxially; involucral bracts absent; Death Valley, California | Gilmania |
15. Perianths glabrous, smooth or minutely pustulose; involucral bracts (3-)4(-7); se California, s Nevada, sw Utah | Johanneshowellia |
16. Involucres not tubular; se California, ec Nevada | Goodmania |
16. Involucres tubular; widespread. [17. Shifted to left margin.—Ed.] | → 17 |
17. Involucral awns 5(-6); tepals 3-lobed or laciniate apically; achenes 3-gonous; s California | Sidotheca |
17. Involucral awns either (3-)4 or 7-36; tepals mostly entire, rarely irregularly divided or retuse apically; achenes globose-lenticular; widespread | → 18 |
18. Involucral awns (3-)4; leaf blades linear to spatulate, margins entire; se Washington e to sw Wyoming and s to Mexico | Oxytheca |
18. Involucral awns 7 or more, rarely 4; leaf blades broadly obovate or spatulate to oblong or oblanceolate, margins ciliate-denticulate; s California | Acanthoscyphus |
|
FNA vol. 5, p. 242. |
FNA vol. 5, p. 218. Author: James L. Reveal. |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
Polygonaceae |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
|
E. microthecum var. alpinum, E. microthecum var. ambiguum, E. microthecum var. arceuthinum, E. microthecum var. johnstonii, E. microthecum var. lacus-ursi, E. microthecum var. laxiflorum, E. microthecum var. microthecum, E. microthecum var. panamintense, E. microthecum var. phoeniceum, E. microthecum var. schoolcraftii, E. microthecum var. simpsonii, E. microthecumvar. corymbosoides |
Acanthoscyphus, Aristocapsa, Centrostegia, Chorizanthe, Dedeckera, Dodecahema, Eriogonum, Gilmania, Goodmania, Hollisteria, Johanneshowellia, Lastarriaea, Mucronea, Nemacaulis, Oxytheca, Pterostegia, Sidotheca, Stenogonum, Systenotheca |
Nuttall: Proc. Acad. Nat. Sci. Philadelphia 4: 15. (1848) |
Arnott: in M. Napier, Encycl. Brit. ed. 7, 5: 126. (1832) |
| |