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slender buckwheat, slender wild buckwheat

spearleaf buckwheat, spearleaf wild buckwheat, spreading wild buckwheat

Habit Subshrubs or shrubs, erect to spreading, not scapose, 0.2–1.5 × (0.6–)1–13(–16) dm, white- to tannish-tomentose, floccose, or glabrous. Shrubs, subshrubs, or herbs, spreading to erect, not scapose, (1–)1.5–5 × 2–5(–8) dm, glabrous or rarely floccose to tomentose, grayish.
Stems

spreading to erect, typically without persistent leaf bases, up to 1/2 height of plant;

caudex stems absent or spreading;

aerial flowering stems erect to spreading, slender, solid, not fistulose, 0.05–1.5 dm, lanate, tomentose, floccose, subglabrous, or glabrous.

spreading or erect, usually without persistent leaf bases, up to 1/2 or more height of plant;

caudex stems absent or matted to spreading;

aerial flowering stems spreading to erect, slender, solid, not fistulose, 0.3–3 dm, glabrous or, rarely, floccose to tomentose, tomentose among leaves.

Leaves

cauline, 1 per node or fasciculate;

petiole 0.1–0.5 cm, tomentose to floccose or glabrous;

blade usually elliptic, sometimes linear to obovate, 0.3–3.5 × (0.07–)0.1–1.2 cm, tomentose abaxially, less so or glabrous adaxially, margins occasionally revolute.

basal or cauline on proximal 1/2 of stem, 1 per node;

petiole 0.5–2 cm, tomentose to floccose or glabrous;

blade narrowly lanceolate or oblanceolate to elliptic, 1.5–7(–9) × 0.2–2 cm, velvety- to densely white-tomentose abaxially, sparsely tomentose to thinly floccose or glabrous and green adaxially, margins plane or occasionally crenulate.

Inflorescences

cymose, compact, often flat-topped, 0.5–6(–12) × 1–10(–13) cm;

branches dichotomous, whitish-lanate to brownish- or reddish-tomentose to floccose or glabrate, infrequently green or gray and subglabrous or glabrous;

bracts 3, scalelike, linear to triangular, 1–5 mm.

cymose, dense to more commonly open, 2–25 × 2–20 cm;

branches dichotomous, glabrous or, rarely, floccose;

bracts 3, scalelike, usually triangular, and 1–3 mm, or occasionally leaflike, 8–30 mm, and otherwise similar to leaf blades.

Peduncles

absent or mostly erect, slender, 0.3–1.5 cm, tomentose to floccose.

absent or erect, 0.1–0.8 cm, glabrous.

Involucres

1 per node, turbinate, (1.5–)2–3.5(–4) × 1.3–2.5(–3) mm, tomentose, floccose, subglabrous, or glabrous;

teeth 5, erect, (0.3–)0.5–1(–1.7) mm.

1 per node or 2–5 per cluster, turbinate to turbinate-campanulate, 2.5–4 × (1.3–)1.5–3.5(–4) mm, glabrous;

teeth 5, erect, 0.4–0.9 mm.

Flowers

1.5–3(–4) mm;

perianth yellow or white to pink, orange, rose, red, or occasionally cream, glabrous;

tepals connate proximal 1/5–2/5, essentially monomorphic, oblong to obovate;

stamens usually exserted, 2.5–4 mm;

filaments sparsely to densely puberulent proximally.

2–3.5(–4) mm;

perianth white, glabrous;

tepals connate proximal 1/4–1/3, monomorphic, oblanceolate, elliptic to oblong or obovate;

stamens exserted, 2–4 mm;

filaments pilose proximally.

Achenes

brown, 1.5–3 mm, glabrous.

light brown to brown, 2–3 mm, glabrous except (typically) for slightly papillate beak.

Eriogonum microthecum

Eriogonum lonchophyllum

Phenology Flowering Jun–Oct.
Habitat Heavy gumbo clay soil or (at higher elevations) sandy-loam to gravelly or rocky soil and outcrops, mixed grassland, saltbush, blackbrush, and sagebrush communities, pinyon-juniper and montane conifer woodlands
Elevation 1400-2900 m (4600-9500 ft)
Distribution
from FNA
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; NM; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 13 (13 in the flora).

Eriogonum microthecum is used as browse by deer and to a lesser degree by cattle and sheep. Some forms are now in cultivation. The species is reportedly used by the Piute of Nevada in the treatment of tuberculosis, lameness, rheumatism, and bladder trouble (P. Train et al. 1941). S. A. Weber and P. D. Seaman (1985) stated that A. F. Whiting found the plants being used as a tea by the Havasupai in northern Arizona. Members of E. microthecum are food plants for subspecies of the rare pallid blue butterfly (Euphilotes pallescens). Also found on this species is the cythera metalmark (Apodemia mormo cythera). Some authors have referred E. effusum to this species, even though the ranges of the two species do not overlap and intermediates are unknown.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eriogonum lonchophyllum is widespread and occasionally rather common in sagebrush-dominated communities in the Rocky Mountains of Colorado, northern New Mexico and eastern Utah. The distribution is fragmented, many of the populations are markedly distinct, and several have been provided with names. The continued recognition of E. coloradense is probably dubious given what is already reduced here to synonymy under E. lonchophyllum. Eriogonum lonchophyllum typically has glabrous flowering stems and inflorescence branches, yet floccose to tomentose individuals are known from Eagle (Reveal & Davidse 861, UTC) and Garfield (Goodrich & D. Nelson 24582, BRY) counties in Colorado. Plants representing the typical expression of the species differ from the majority of populations assigned here in having broad leaves and a sprawling habit. Such plants grow in deep shade under tall conifers and are known currently only from the type locality. Variation in the size, shape, and distribution of leaves has been used to distinguish varieties of E. lonchophyllum. At their extremes, the differences are remarkable. For example, plants with linear, basal leaves have been named var. nudicaule. Such plants are found in the New Mexico counties of Rio Arriba, Sandoval, and Santa Fe. Another herbaceous phase also with basal, but broader leaves and known from the Book Cliffs and Tavaputs Plateau regions of northeastern Emery, southern Uintah, and northern Grand counties, Utah, and in Rio Blanco County, Colorado, has been named var. intermontanum. The low, spreading subshrubby phase found along the foothills of the Front Ranges in south-central Colorado (El Paso, Fremont, Jefferson, Las Animas, Otero, and Pueblo counties) and adjacent north-central New Mexico (Colfax and Taos counties) is known as var. fendlerianum. These plants have velvety, tomentose leaf blades. Another shrubby phase is found in and around the Dinosaur National Park in Daggett and Uintah counties, Utah, and in northwestern Rio Blanco County, Colorado. This is var. saurinum, characterized by having sheathing leaf blades along the flowering stems that vary from narrowly oblanceolate to narrowly elliptic; it is occasionally confused with E. corymbosum in that area. At the north end of Mesa Verde National Park and on Sleeping Ute Mountain of Montezuma County, Colorado, is a large shrub that outwardly resembles E. corymbosum in size and shape. It is similar to, but larger than, the more eastern var. fendlerianum. The most common phase of the species (as represented by var. humivagans, including E. salicinum, E. scoparium, and E. tristichum) is found mainly on fine, clayey to gravelly, alluvial soils in the Rocky Mountains and western desert ranges of Colorado (Archuleta, Delta, Dolores, Eagle, Garfield, Gunnison, La Plata, Mesa, Moffat, Montezuma, Montrose, Ouray, Rio Blanco, Saguache, and San Miguel counties), northern New Mexico (Rio Arriba, Sandoval, and San Juan counties), and San Juan County, Utah. The majority of the plants have long, narrow leaf blades (2–5 mm wide), but populations with broader leaves are common in southwestern Colorado. It is this latter expression of E. lonchophyllum that can be difficult to distinguish from E. corymbosum in portions of eastern Utah and western Colorado. A single collection (Rammel s.n., 1872, MIN, US) was supposedly gathered in Texas. Until this is confirmed, the species is not considered to be part of the Texas flora.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Perianths yellow
→ 2
1. Perianths various shades of white, cream, orange, pink, or red, not yellow
→ 4
2. Flowering stems and inflorescence branches usually glabrous; e Oregon and wc Idaho
var. microthecum
2. Flowering stems and inflorescence branches tomentose to floccose; se Oregon and sw Idaho s to e California and w Nevada
→ 3
3. Leaf blades (0.2-)0.3-0.6(-0.8) cm wide; flowers (1.5-)2-2.5(-3) mm; involucres 2-2.5 mm; achenes 1.5-2 mm; se Oregon and sw Idaho s to e California and w Nevada
var. ambiguum
3. Leaf blades 0.5-1.2 cm wide; flowers 2.5-3 mm; involucres 2.5-4 mm; achenes 2.5-3 mm; ne California, nw Nevada
var. schoolcraftii
4. Tomentum whitish (see also var. alpinum of the Sierra Nevada, California); flowering stems and inflorescence branches infrequently glabrous
→ 5
4. Tomentum brownish or reddish (may be white in var. alpinum), or flowering stems and inflorescence branches essentially glabrous
→ 6
5. Leaf margins not revolute; flowering inflorescence branches floccose or glabrous; northern phase of species
var. laxiflorum
5. Leaf margins revolute or nearly so; flowering inflorescence branches lanate to tomentose, or if subglabrous or glabrous, then southern phase of species
var. simpsonii
6. Plants shrubs, 3-6 dm
→ 7
6. Plants subshrubs, 0.2-1.5(-2) dm
→ 8
7. Flowering stems and inflorescence branches tomentose when young, becoming floccose at maturity; flowers 1.5-2(-2.5) mm; achenes 1.8-2 mm; Death Valley region, California
var. panamintense
7. Flowering stems and inflorescence branches lanate to tomentose at maturity; flowers 2-2.5(-3) mm; achenes 2.5-3 mm; Transverse Ranges, California
E. microthecumvar. corymbosoides
8. Leaf blades elliptic or ovate, margins not revolute; flowers (1.5-)2-3.5(-4) mm
→ 9
8. Leaf blades linear or linear-oblanceolate to narrowly elliptic, margins often revolute; flowers 1.5-2.5 mm
→ 10
9. Leaf blades 0.5-1 × (0.2-)0.3-0.5(-0.6) cm; involucres (2-)2.5-3 mm; flowers (2.5-)3-3.5(-4) mm; San Gabriel Mountains, California
var. johnstonii
9. Leaf blades 0.3-0.7(-0.8) × 0.1-0.4 cm; involucres 2.5-3.5 mm; flowers (1.5-)2-3.5 mm; desert ranges of se California, c Nevada, and Utah1f. Eriogonum microthecum var. lapidicola [8. Shifted to left margin—Ed.]
→ 8
10. Involucres 3-4 mm; perianths cream; San Bernardino Mountains, California
var. lacus-ursi
10. Involucres (1.5-)2-3 mm; perianths white, pink, red, or rose; Sierra Nevada of California, or e Nevada and w Utah
→ 11
11. Flowering stems glabrous; leaf blades sparsely floccose or glabrous adaxially; desert ranges, se Nevada
var. arceuthinum
11. Flowering stems white- to brownish-floccose to subglabrous, or reddish-tomentose to floccose; leaf blades floccose to subglabrous adaxially; California or wc Utah
→ 12
12. Tomentum white to brownish; Sierra Nevada, California
var. alpinum
12. Tomentum reddish; desert ranges, wc Utah
var. phoeniceum
Source FNA vol. 5, p. 242. FNA vol. 5, p. 262.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. microthecum var. alpinum, E. microthecum var. ambiguum, E. microthecum var. arceuthinum, E. microthecum var. johnstonii, E. microthecum var. lacus-ursi, E. microthecum var. laxiflorum, E. microthecum var. microthecum, E. microthecum var. panamintense, E. microthecum var. phoeniceum, E. microthecum var. schoolcraftii, E. microthecum var. simpsonii, E. microthecumvar. corymbosoides
Synonyms E. ainsliei, E. corymbosum var. humivagans, E. effusum subsp. ainsliei, E. effusum subsp. fendlerianum, E. effusum var. nudicaule, E. effusum subsp. salicinum, E. fendlerianum, E. humivagans, E. intermontanum, E. lonchophyllum var. fendlerianum, E. lonchophyllum var. humivagans, E. lonchophyllum var. intermontanum, E. lonchophyllum var. nudicaule, E. lonchophyllum var. saurinum, E. nudicaule, E. nudicaule subsp. scoparium, E. nudicaule subsp. tristichum, E. salicinum, E. saurinum, E. scoparium, E. tristichum
Name authority Nuttall: Proc. Acad. Nat. Sci. Philadelphia 4: 15. (1848) Torrey & A. Gray: Proc. Amer. Acad. Arts 8: 173. (1870)
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