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marum-leaf buckwheat, marum-leaf wild buckwheat, mountain buckwheat

Thurber eriogonum, Thurber's buckwheat, Thurber's wild buckwheat

Habit Herbs, matted, dioecious, (0.1–)0.5–5 × 2–8 dm, floccose or glabrous. Herbs, spreading, annual, 0.5–4 dm, glabrate, glabrous or sparsely glandular, greenish, grayish, or reddish.
Stems

caudex spreading;

aerial flowering stems erect, slender, solid, not fistulose, arising at nodes of caudex branches and at distal nodes of short, non-flowering aerial branches, (0.1–)0.5–4 dm, floccose or glabrous.

caudex absent;

aerial flowering stems erect, solid, not fistulose, 0.3–1 dm, often sparsely tomentose and glandular proximally.

Leaves

in loose to compact basal rosettes;

petiole 0.3–2 cm, often tomentose;

blade ovate to oval, 0.3–3 × 0.3–1 cm, densely tannish- to brownish-lanate abaxially, glabrate and green to olive green adaxially, margins entire, plane.

basal;

petiole 1–3 cm;

blade oblong to narrowly ovate, 0.8–4.5 × 0.5–3 cm, densely white-tomentose abaxially, floccose or glabrous and greenish adaxially, margins often crenulate.

Inflorescences

capitate, 0.5–2 cm wide, mature pistillate plants open and elongate-umbellate, 1–5 × 1–7 cm;

branches usually glabrous;

bracts 5–10, leaflike, 0.2–0.8 cm, often absent immediately below involucre.

cymose, mostly diffuse, 5–30 × 5–50 cm;

branches sparsely glandular to glabrate or glabrous;

bracts 3, scalelike, 1–2.5 × 1–2.5 mm.

Peduncles

erect, straight, capillary, 0.5–2.5 cm, glabrous and glandular-puberulent distally.

Involucres

1 per node but occasionally appearing congested, turbinate and 2–3 × 1.5–2 mm or campanulate and 4–5 × 5–7 mm, sparsely tomentose or glabrous;

teeth 5–6, erect, 0.4–0.7 mm.

broadly turbinate, 1.8–2 × 1.8–2 mm, minutely glandular-puberulent;

teeth 5, erect, 0.4–0.6 mm.

Flowers

stipelike base 0.5–1 mm;

perianth yellow, glabrous; staminate flowers 1.5–3 mm, tepals ovate; pistillate flowers 4–7 mm, tepals oblanceolate, often becoming reddish in fruit;

stamens exserted, 2–3 mm;

filaments pilose proximally.

1–1.7 mm;

perianth white with greenish or reddish midribs, becoming red, glandular-puberulent with a tuft of long white hairs adaxially;

tepals dimorphic, those of outer whorl broadly pandurate or flabellate, those of inner whorl oblanceolate;

stamens included, 0.7–1.2 mm;

filaments mostly glabrous.

Achenes

light brown to brown, 3.5–5 mm, glabrous except for sparsely pubescent beak.

brown to black, usually lenticular, 0.6–0.8 mm, glabrous.

2n

= 32.

= 40.

Eriogonum marifolium

Eriogonum thurberi

Phenology Flowering year-round.
Habitat Sandy flats, washes, and slopes, saltbush, greasewood, and creosote bush communities, oak, pinyon and/or juniper woodlands, (montane conifer woodlands in Mexico)
Elevation 100-1200 m (300-3900 ft)
Distribution
from FNA
CA; NV; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NM; Mexico (Baja California, Sonora)
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Eriogonum marifolium is dioecious, with the two sexes having such markedly different morphology that collectors often consider each to represent a different species. In Oregon, male plants of E. marifolium and E. umbellatum var. haussknechtii can be difficult to distinguish. In El Dorado and Placer counties, California, herbarium specimens of E. marifolium and E. incanum can be difficult to separate, as their distribution ranges overlap there.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eriogonum thurberi is common to abundant but rarely weedy in northwestern Mexico, southern California (Inyo, Los Angeles, Orange, Riverside, San Bernardino, and San Diego counties), and southern Arizona (Cochise, Gila, Graham, Maricopa, Mohave, Pima, Pinal, Santa Cruz, and Yavapai counties). It just enters New Mexico (Grant County). It and E. thomasii are the annual members of the E. cernuum complex typically found on the Mojave and Sonoran deserts, with E. thurberi extending farther to the east and E. thomasii farther to the north. The two occasionally grow together.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Involucres turbinate, 2-3 × 1.5-2 mm; ne California, wc Nevada, s Oregon, disjunct to Yakima County, Washington
var. marifolium
1. Involucres campanulate, 4-5 × 5-7 mm; nc California
var. cupulatum
Source FNA vol. 5, p. 365. FNA vol. 5, p. 403.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Oligogonum Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. marifolium var. cupulatum, E. marifolium var. marifolium
Synonyms E. cupulatum, E. marifolium var. apertum E. cernuum subsp. thurberi, E. cernuum subsp. viscosum
Name authority Torrey & A. Gray: Proc. Amer. Acad. Arts 8: 161. (1870) Torrey: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 176. (1859)
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