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marum-leaf buckwheat, marum-leaf wild buckwheat, mountain buckwheat

annual buckwheat, annual wild buckwheat

Habit Herbs, matted, dioecious, (0.1–)0.5–5 × 2–8 dm, floccose or glabrous. Herbs, 5–20 × 5–10 dm, grayish.
Stems

caudex spreading;

aerial flowering stems erect, slender, solid, not fistulose, arising at nodes of caudex branches and at distal nodes of short, non-flowering aerial branches, (0.1–)0.5–4 dm, floccose or glabrous.

Aerial flowering stems

slender, 4–10(–15) dm, floccose to densely tomentose.

Leaves

in loose to compact basal rosettes;

petiole 0.3–2 cm, often tomentose;

blade ovate to oval, 0.3–3 × 0.3–1 cm, densely tannish- to brownish-lanate abaxially, glabrate and green to olive green adaxially, margins entire, plane.

petiole (rosette) 0.3–1.2 cm, or petiole (cauline) 0.2–0.5 cm, tomentose to floccose;

blade oblanceolate to oblong, 1–7 × 0.3–1.5 cm, densely tomentose abaxially, floccose adaxially not thickened and auriculate-subclasping proximally;

margins entire or slightly revolute.

Inflorescences

capitate, 0.5–2 cm wide, mature pistillate plants open and elongate-umbellate, 1–5 × 1–7 cm;

branches usually glabrous;

bracts 5–10, leaflike, 0.2–0.8 cm, often absent immediately below involucre.

3–10 × 2–7 cm;

bracts triangular, 1–4 mm.

Peduncles

0.1–0.5 cm, tomentose to floccose.

Involucres

1 per node but occasionally appearing congested, turbinate and 2–3 × 1.5–2 mm or campanulate and 4–5 × 5–7 mm, sparsely tomentose or glabrous;

teeth 5–6, erect, 0.4–0.7 mm.

turbinate to campanulate, 2.5–4 × 2–3 mm, tomentose to floccose abaxially, glabrous adaxially;

teeth 5–6, 0.4–1 mm.

Flowers

stipelike base 0.5–1 mm;

perianth yellow, glabrous; staminate flowers 1.5–3 mm, tepals ovate; pistillate flowers 4–7 mm, tepals oblanceolate, often becoming reddish in fruit;

stamens exserted, 2–3 mm;

filaments pilose proximally.

1–2.5 mm;

perianth white or cream to rose;

tepals: those of outer whorl obovate, 1–2 × 0.9–1.5 mm, those of inner whorl narrowly ovate to oblong, 1.5–4 × 1.2–1.8 mm;

stamens usually included, 1–2 mm.

Achenes

light brown to brown, 3.5–5 mm, glabrous except for sparsely pubescent beak.

1.5–2 mm.

2n

= 32.

= 40.

Eriogonum marifolium

Eriogonum annuum

Phenology Flowering Apr–Nov.
Habitat Sandy flats, slopes, dunes, and banks, mixed grassland, oak and conifer woodlands
Elevation (0-)100-1900(-2300) m ((0-)300-6200(-7500) ft)
Distribution
from FNA
CA; NV; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; KS; MT; ND; NE; NM; OK; SD; TX; WY; Mexico (Chihuahua)
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Eriogonum marifolium is dioecious, with the two sexes having such markedly different morphology that collectors often consider each to represent a different species. In Oregon, male plants of E. marifolium and E. umbellatum var. haussknechtii can be difficult to distinguish. In El Dorado and Placer counties, California, herbarium specimens of E. marifolium and E. incanum can be difficult to separate, as their distribution ranges overlap there.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eriogonum annuum is widespread and common to locally abundant or even weedy on the Great Plains of the central United States and extreme north-central Mexico. It was collected in Sherburne County, Minnesota, in 1982, but that population did not persist. The species was recently found as an introduction at Sandy Hook in Monmouth County, New Jersey (Snyder & McArthur s.n., NY), but its fate there remains to be determined. Unfortunately, this weedy species recently has been introduced into northern Arizona as a roadside wild flower. C. L. Perez et al. (1998) have demonstrated that the seed bank can be rich in seeds of this species, but germination rates are low.

The Lakota people traditionally used the annual wild buckwheat as an aid in the treatment of sore mouths in children, seemingly in association with teething (D. J. Rogers 1980). Leaves were used to stain buffalo and deer hides by the Kiowa (P. A. Vestal and R. E. Schultes 1939). Vestal (1952) stated that the species was considered a “life medicine” by the Navajo (Diné) people; it was used also for protection against witches. It is likely that E. annuum was obtained by the Navajo through trade, but it might have been grown locally in historic times where the species recently has been reintroduced.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Involucres turbinate, 2-3 × 1.5-2 mm; ne California, wc Nevada, s Oregon, disjunct to Yakima County, Washington
var. marifolium
1. Involucres campanulate, 4-5 × 5-7 mm; nc California
var. cupulatum
Source FNA vol. 5, p. 365. FNA vol. 5.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Oligogonum Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Micrantha
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. marifolium var. cupulatum, E. marifolium var. marifolium
Synonyms E. cupulatum, E. marifolium var. apertum E. annuum subsp. cymosum, E. annuum subsp. hitchcockii
Name authority Torrey & A. Gray: Proc. Amer. Acad. Arts 8: 161. (1870) Nuttall: Trans. Amer. Philos. Soc., n. s. 5: 164. (1835)
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