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angle stermed buckwheat, spotted buckwheat, spotted wild buckwheat
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Blue Mountain buckwheat, strict buckwheat
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| Herbs, erect to spreading, annual, 1–2(–3) dm, tomentose, greenish to reddish. |
Herbs, erect, loosely to densely matted, not scapose, 1–5 × 1–10 dm, tomentose or glabrous. |
caudex absent; aerial flowering stems erect, not striated or angled, solid, not fistulose, 0.1–0.5 dm, tomentose. |
spreading to erect, with or without persistent leaf bases, up to 1/4 height of plant; caudex stems absent or spreading to matted; aerial flowering stems erect to spreading, slender, solid, not fistulose, 1–3 dm, tomentose or glabrous. |
basal and cauline; basal: petiole 0.3–1 cm, floccose, blade lanceolate to obovate, 1–3(–4) × 1–1.5(–2) cm, tomentose abaxially, sparsely floccose to glabrate and grayish to greenish adaxially, margins entire or infrequently crenulate; cauline sessile, blade lanceolate to oblanceolate, 0.5–2 × 0.3–1 cm, similar to basal blade. |
basal, 1 per node; petiole rarely twisted or curled, 1–6 cm, mostly tomentose; blade elliptic to ovate, 0.5–2.5(–4) × (0.3–)0.5–1.5 cm, lanate, tomentose to floccose on both surfaces, sometimes sparsely tomentose to floccose and greenish or floccose to subglabrous or glabrous adaxially, margins plane. |
cymose, open, 5–25 × 10–30 cm; branches tomentose; bracts 3, scalelike, 0.5–2.5 × 1–2 mm. |
umbellate-cymose to cymose, 1–20 × 3–25 cm; branches dichotomous, tomentose to floccose or less often glabrous; bracts 3, scalelike, triangular, 1–3 mm. |
spreading, straight or nearly so, filiform, (0.5–)1–3 cm, glandular-puberulent. |
absent. |
campanulate, 1–1.5(–2) × 1.5–3(–3.5) mm, glandular-puberulent; teeth 5, erect, 0.4–0.8 mm. |
1 per node, rarely 2–5 per cluster, narrowly turbinate to turbinate-campanulate, 4–6 × 1.5–5 mm, tomentose or glabrous; teeth 5, erect, 0.5–1.3 mm. |
1–2.5 mm; perianth white to yellow, becoming pink or red, with a conspicuous rose-purple spot on each outer tepal, glandular-puberulent; tepals dimorphic, those of outer whorl elliptic to roundish or obovate, inflated from base to middle, those of inner whorl lanceolate; stamens included, 1–1.5(–2) mm; filaments pilose proximally. |
3–5(–6) mm; perianth yellow or white to rose or purple, glabrous; tepals connate proximally, dimorphic, those of outer whorl elliptic to nearly orbiculate, 2–3 × 2–3 mm, those of inner whorl oblanceolate to oblong, 3–4 × 1–2 mm; stamens included to slightly exserted, 2–5 mm; filaments pilose proximally. |
brown, 3-gonous, 1–1.5 mm, glabrous. |
light brown to brown, 3–3.5 mm, glabrous. |
= 40. |
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| Flowering Apr–Nov. |
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| Sandy to gravelly or clayey flats and slopes, mixed grassland, saltbush, creosote bush, and sagebrush communities, pinyon-juniper and montane conifer woodlands |
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| 100-2500 m (300-8200 ft) |
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AZ; CA; ID; NM; NV; OR; UT; WA; Mexico (Baja California)
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CA; ID; MT; NV; OR; WA
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Eriogonum maculatum is the most common and widespread expression of the E. angulosum complex, being found in Arizona, California, southwestern Idaho, Nevada, northwestern New Mexico, eastern Oregon, western Utah, and southeastern Washington. It is often common to abundant and may even be weedy, especially along roadsides intermixed with other annual wild buckwheats. The swollen bases of the outer tepals readily distinguish the species. The greatest concentrations of the spotted wild buckwheat are found in the Mojave Desert and the Great Basin. A specimen supposedly found on the “Laramie Plains” of Wyoming (Parry s.n., 1882, ISC) is discounted as to location. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 4 (4 in the flora). Eriogonum strictum, E. niveum, and E. ovalifolium form a complex of closely related species differing in leaf, inflorescence branching, and flower features. Variety proliferum appears to be the basal entity of the complex, approaching both E. niveum and E. ovalifolium var. pansum in its pubescence and branching pattern. Also, specimens of var. proliferum are sometimes difficult to differentiate from E. nudum var. oblongifolium. Careful observation, though, will permit well-made collections to be easily distributed among the individual species. An alternative taxonomy is to reduce all of the taxa to E. ovalifolium and recognize a series of subspecies and varieties. It is possible that additional study will show that E. strictum is sufficiently distinct from its tomentose to floccose counterparts to justify recognition of E. proliferum. In that case, both var. anserinum and var. greenei would be assigned to the latter species. Or, one could follow C. L. Hitchcock et al. (1955–1969, vol. 2) and recognize subsp. strictum as distinct from subsp. proliferum, with the latter consisting of varieties proliferum, anserinum, and greenei. Members of the Eriogonum strictum are food plants for the Bauer’s dotted-blue butterfly (Euphilotes baueri). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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1. Inflorescence branches glabrous | var. strictum |
1. Inflorescence branches tomentose to floccose | → 2 |
| var. anserinum |
2. Perianths white to rose or purple | → 3 |
3. Leaf blades grayish-tomentose to floccose on both surfaces, or greenish-tomentose to floccose adaxially | var. proliferum |
3. Leaf blades densely whitish-lanate to tomentose on both surfaces | var. greenei |
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| FNA vol. 5, p. 411. |
FNA vol. 5, p. 317. |
| Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
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| E. angulosum subsp. maculatum, E. angulosum var. maculatum, E. angulosum var. rectipes |
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| A. Heller: Muhlenbergia 2: 188. (1906) |
Bentham: Trans. Linn. Soc. London 17: 414. (1836) |
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CA, 
OR, 
WA
Flora NW, 
PNW Herbaria
WildflowerSearch
iNaturalist (observations)
USDA Plants Database
