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angle stermed buckwheat, spotted buckwheat, spotted wild buckwheat
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spearleaf buckwheat, spearleaf wild buckwheat, spreading wild buckwheat
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| Herbs, erect to spreading, annual, 1–2(–3) dm, tomentose, greenish to reddish. |
Shrubs, subshrubs, or herbs, spreading to erect, not scapose, (1–)1.5–5 × 2–5(–8) dm, glabrous or rarely floccose to tomentose, grayish. |
caudex absent; aerial flowering stems erect, not striated or angled, solid, not fistulose, 0.1–0.5 dm, tomentose. |
spreading or erect, usually without persistent leaf bases, up to 1/2 or more height of plant; caudex stems absent or matted to spreading; aerial flowering stems spreading to erect, slender, solid, not fistulose, 0.3–3 dm, glabrous or, rarely, floccose to tomentose, tomentose among leaves. |
basal and cauline; basal: petiole 0.3–1 cm, floccose, blade lanceolate to obovate, 1–3(–4) × 1–1.5(–2) cm, tomentose abaxially, sparsely floccose to glabrate and grayish to greenish adaxially, margins entire or infrequently crenulate; cauline sessile, blade lanceolate to oblanceolate, 0.5–2 × 0.3–1 cm, similar to basal blade. |
basal or cauline on proximal 1/2 of stem, 1 per node; petiole 0.5–2 cm, tomentose to floccose or glabrous; blade narrowly lanceolate or oblanceolate to elliptic, 1.5–7(–9) × 0.2–2 cm, velvety- to densely white-tomentose abaxially, sparsely tomentose to thinly floccose or glabrous and green adaxially, margins plane or occasionally crenulate. |
cymose, open, 5–25 × 10–30 cm; branches tomentose; bracts 3, scalelike, 0.5–2.5 × 1–2 mm. |
cymose, dense to more commonly open, 2–25 × 2–20 cm; branches dichotomous, glabrous or, rarely, floccose; bracts 3, scalelike, usually triangular, and 1–3 mm, or occasionally leaflike, 8–30 mm, and otherwise similar to leaf blades. |
spreading, straight or nearly so, filiform, (0.5–)1–3 cm, glandular-puberulent. |
absent or erect, 0.1–0.8 cm, glabrous. |
campanulate, 1–1.5(–2) × 1.5–3(–3.5) mm, glandular-puberulent; teeth 5, erect, 0.4–0.8 mm. |
1 per node or 2–5 per cluster, turbinate to turbinate-campanulate, 2.5–4 × (1.3–)1.5–3.5(–4) mm, glabrous; teeth 5, erect, 0.4–0.9 mm. |
1–2.5 mm; perianth white to yellow, becoming pink or red, with a conspicuous rose-purple spot on each outer tepal, glandular-puberulent; tepals dimorphic, those of outer whorl elliptic to roundish or obovate, inflated from base to middle, those of inner whorl lanceolate; stamens included, 1–1.5(–2) mm; filaments pilose proximally. |
2–3.5(–4) mm; perianth white, glabrous; tepals connate proximal 1/4–1/3, monomorphic, oblanceolate, elliptic to oblong or obovate; stamens exserted, 2–4 mm; filaments pilose proximally. |
brown, 3-gonous, 1–1.5 mm, glabrous. |
light brown to brown, 2–3 mm, glabrous except (typically) for slightly papillate beak. |
= 40. |
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| Flowering Apr–Nov. |
Flowering Jun–Oct. |
| Sandy to gravelly or clayey flats and slopes, mixed grassland, saltbush, creosote bush, and sagebrush communities, pinyon-juniper and montane conifer woodlands |
Heavy gumbo clay soil or (at higher elevations) sandy-loam to gravelly or rocky soil and outcrops, mixed grassland, saltbush, blackbrush, and sagebrush communities, pinyon-juniper and montane conifer woodlands |
| 100-2500 m (300-8200 ft) |
1400-2900 m (4600-9500 ft) |
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AZ; CA; ID; NM; NV; OR; UT; WA; Mexico (Baja California)
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CO; NM; UT
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Eriogonum maculatum is the most common and widespread expression of the E. angulosum complex, being found in Arizona, California, southwestern Idaho, Nevada, northwestern New Mexico, eastern Oregon, western Utah, and southeastern Washington. It is often common to abundant and may even be weedy, especially along roadsides intermixed with other annual wild buckwheats. The swollen bases of the outer tepals readily distinguish the species. The greatest concentrations of the spotted wild buckwheat are found in the Mojave Desert and the Great Basin. A specimen supposedly found on the “Laramie Plains” of Wyoming (Parry s.n., 1882, ISC) is discounted as to location. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eriogonum lonchophyllum is widespread and occasionally rather common in sagebrush-dominated communities in the Rocky Mountains of Colorado, northern New Mexico and eastern Utah. The distribution is fragmented, many of the populations are markedly distinct, and several have been provided with names. The continued recognition of E. coloradense is probably dubious given what is already reduced here to synonymy under E. lonchophyllum. Eriogonum lonchophyllum typically has glabrous flowering stems and inflorescence branches, yet floccose to tomentose individuals are known from Eagle (Reveal & Davidse 861, UTC) and Garfield (Goodrich & D. Nelson 24582, BRY) counties in Colorado. Plants representing the typical expression of the species differ from the majority of populations assigned here in having broad leaves and a sprawling habit. Such plants grow in deep shade under tall conifers and are known currently only from the type locality. Variation in the size, shape, and distribution of leaves has been used to distinguish varieties of E. lonchophyllum. At their extremes, the differences are remarkable. For example, plants with linear, basal leaves have been named var. nudicaule. Such plants are found in the New Mexico counties of Rio Arriba, Sandoval, and Santa Fe. Another herbaceous phase also with basal, but broader leaves and known from the Book Cliffs and Tavaputs Plateau regions of northeastern Emery, southern Uintah, and northern Grand counties, Utah, and in Rio Blanco County, Colorado, has been named var. intermontanum. The low, spreading subshrubby phase found along the foothills of the Front Ranges in south-central Colorado (El Paso, Fremont, Jefferson, Las Animas, Otero, and Pueblo counties) and adjacent north-central New Mexico (Colfax and Taos counties) is known as var. fendlerianum. These plants have velvety, tomentose leaf blades. Another shrubby phase is found in and around the Dinosaur National Park in Daggett and Uintah counties, Utah, and in northwestern Rio Blanco County, Colorado. This is var. saurinum, characterized by having sheathing leaf blades along the flowering stems that vary from narrowly oblanceolate to narrowly elliptic; it is occasionally confused with E. corymbosum in that area. At the north end of Mesa Verde National Park and on Sleeping Ute Mountain of Montezuma County, Colorado, is a large shrub that outwardly resembles E. corymbosum in size and shape. It is similar to, but larger than, the more eastern var. fendlerianum. The most common phase of the species (as represented by var. humivagans, including E. salicinum, E. scoparium, and E. tristichum) is found mainly on fine, clayey to gravelly, alluvial soils in the Rocky Mountains and western desert ranges of Colorado (Archuleta, Delta, Dolores, Eagle, Garfield, Gunnison, La Plata, Mesa, Moffat, Montezuma, Montrose, Ouray, Rio Blanco, Saguache, and San Miguel counties), northern New Mexico (Rio Arriba, Sandoval, and San Juan counties), and San Juan County, Utah. The majority of the plants have long, narrow leaf blades (2–5 mm wide), but populations with broader leaves are common in southwestern Colorado. It is this latter expression of E. lonchophyllum that can be difficult to distinguish from E. corymbosum in portions of eastern Utah and western Colorado. A single collection (Rammel s.n., 1872, MIN, US) was supposedly gathered in Texas. Until this is confirmed, the species is not considered to be part of the Texas flora. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 5, p. 411. |
FNA vol. 5, p. 262. |
| Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
| E. angulosum subsp. maculatum, E. angulosum var. maculatum, E. angulosum var. rectipes |
E. ainsliei, E. corymbosum var. humivagans, E. effusum subsp. ainsliei, E. effusum subsp. fendlerianum, E. effusum var. nudicaule, E. effusum subsp. salicinum, E. fendlerianum, E. humivagans, E. intermontanum, E. lonchophyllum var. fendlerianum, E. lonchophyllum var. humivagans, E. lonchophyllum var. intermontanum, E. lonchophyllum var. nudicaule, E. lonchophyllum var. saurinum, E. nudicaule, E. nudicaule subsp. scoparium, E. nudicaule subsp. tristichum, E. salicinum, E. saurinum, E. scoparium, E. tristichum |
| A. Heller: Muhlenbergia 2: 188. (1906) |
Torrey & A. Gray: Proc. Amer. Acad. Arts 8: 173. (1870) |
| |
CA, 
OR, 
WA
Flora NW, 
PNW Herbaria
WildflowerSearch
iNaturalist (observations)
USDA Plants Database
SW CO Wildflowers
