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angle stermed buckwheat, spotted buckwheat, spotted wild buckwheat

Kennedy's buckwheat, Kennedy's wild buckwheat

Habit Herbs, erect to spreading, annual, 1–2(–3) dm, tomentose, greenish to reddish. Herbs, scapose, matted, 0.4–1.5 × 1–4 dm, glabrous or sparsely floccose to tomentose, grayish or reddish.
Stems

caudex absent;

aerial flowering stems erect, not striated or angled, solid, not fistulose, 0.1–0.5 dm, tomentose.

matted, occasionally with persistent leaf bases, up to 1/5 height of plant;

caudex stems matted;

aerial flowering stems scapelike,erect or nearly so, slender, solid, not fistulose, (0.5–)1.5–4.5(–5) dm, glabrous or sparsely floccose to tomentose.

Leaves

basal and cauline;

basal: petiole 0.3–1 cm, floccose, blade lanceolate to obovate, 1–3(–4) × 1–1.5(–2) cm, tomentose abaxially, sparsely floccose to glabrate and grayish to greenish adaxially, margins entire or infrequently crenulate;

cauline sessile, blade lanceolate to oblanceolate, 0.5–2 × 0.3–1 cm, similar to basal blade.

basal, fasciculate in terminal tufts, sometimes 1 per node and sheathing up stem 1–2 cm;

petiole 0.05–1 cm, tomentose;

blade oblanceolate, elliptic, or oblong, 0.2–1(–1.2) × 0.05–0.4 cm, grayish-, brownish-, white-, or reddish-white-tomentose, usually on both surfaces, margins plane or revolute.

Inflorescences

cymose, open, 5–25 × 10–30 cm;

branches tomentose;

bracts 3, scalelike, 0.5–2.5 × 1–2 mm.

capitate, 0.4–1 cm;

branches absent;

bracts 4–7, scalelike, triangular, 0.5–2.5(–3) mm.

Peduncles

spreading, straight or nearly so, filiform, (0.5–)1–3 cm, glandular-puberulent.

absent.

Involucres

campanulate, 1–1.5(–2) × 1.5–3(–3.5) mm, glandular-puberulent;

teeth 5, erect, 0.4–0.8 mm.

3–7 per cluster, turbinate to turbinate-campanulate, 1.5–4 × 1–3.5 mm, rigid, glabrous or tomentose;

teeth 5, erect to spreading, 0.4–1.5 mm.

Flowers

1–2.5 mm;

perianth white to yellow, becoming pink or red, with a conspicuous rose-purple spot on each outer tepal, glandular-puberulent;

tepals dimorphic, those of outer whorl elliptic to roundish or obovate, inflated from base to middle, those of inner whorl lanceolate;

stamens included, 1–1.5(–2) mm;

filaments pilose proximally.

1.5–4 mm, glabrous;

perianth white to pink or rose;

tepals connate proximal 1/4, monomorphic, oblanceolate to elliptic or obovate to nearly oval;

stamens exserted, 1.5–4 mm;

filaments glabrous or sparsely pubescent proximally.

Achenes

brown, 3-gonous, 1–1.5 mm, glabrous.

light brown to brown, 1.8–4 mm, glabrous.

2n

= 40.

Eriogonum maculatum

Eriogonum kennedyi

Phenology Flowering Apr–Nov.
Habitat Sandy to gravelly or clayey flats and slopes, mixed grassland, saltbush, creosote bush, and sagebrush communities, pinyon-juniper and montane conifer woodlands
Elevation 100-2500 m (300-8200 ft)
Distribution
from FNA
AZ; CA; ID; NM; NV; OR; UT; WA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV
[WildflowerSearch map]
[BONAP county map]
Discussion

Eriogonum maculatum is the most common and widespread expression of the E. angulosum complex, being found in Arizona, California, southwestern Idaho, Nevada, northwestern New Mexico, eastern Oregon, western Utah, and southeastern Washington. It is often common to abundant and may even be weedy, especially along roadsides intermixed with other annual wild buckwheats. The swollen bases of the outer tepals readily distinguish the species. The greatest concentrations of the spotted wild buckwheat are found in the Mojave Desert and the Great Basin. A specimen supposedly found on the “Laramie Plains” of Wyoming (Parry s.n., 1882, ISC) is discounted as to location.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 5 (5 in the flora).

The capitate inflorescence and basal leaves of Eriogonum kennedyi distinguish it from the closely related E. wrightii. At higher elevations in the Transverse Ranges of California, the distinction between E. kennedyi var. kennedyi and E. wrightii var. subscaposum is not always clear, especially near the summit of Mt. Pinos. At lower elevations in the San Bernardino Mountains, in more arid habitats, the distinction between E. kennedyi var. austromontanum and E. wrightii var. subscaposum also is blurred. For the most part this intergradation seems to be associated with a reduction in the length of the inflorescence in depauperate expressions of var. subscaposum and not a result of hybridization. The extreme reduction seen in E. wrightii var. olanchense exemplifies the general trend in the California expressions of that species toward depauperate forms. It is felt that E. kennedyi itself is an established segregate that owes its origin to E. wrightii. Members of the species are food plants for Bauer’s dotted-blue butterfly (Euphilotes baueri) and the Ord Mountain metalmark butterfly (Apodemia mormo dialeuca).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf tomentum white; plants forming dense mats; desert ranges, ec California, wc Nevada
var. purpusii
1. Leaf tomentum grayish or reddish- to brownish- white; plants usually forming loose, open mats; southern Sierra Nevada and Transverse Ranges of s California
→ 2
2. Leaf blades oblong, 0.3-0.5 × 0.1-0.4 cm
var. pinicola
2. Leaf blades oblanceolate to elliptic, 0.2-1.2 × 0.05-0.2 cm
→ 3
3. Scapes 0.5-2(-3) cm
var. alpigenum
3. Scapes 4-15 cm
→ 4
4. Leaf blades 0.2-0.4(-0.5) × 0.05-0.15(-0.2) cm; scapes glabrous; involucres 1.5-2.5 mm
var. kennedyi
4. Leaf blades (0.4-)0.6-1(-1.2) × 0.1-0.2 mm; scapes sparsely tomentose to floccose; involucres 2.5-4 mm
var. austromontanum
Source FNA vol. 5, p. 411. FNA vol. 5, p. 292.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. kennedyi var. alpigenum, E. kennedyi var. austromontanum, E. kennedyi var. kennedyi, E. kennedyi var. pinicola, E. kennedyi var. purpusii
Synonyms E. angulosum subsp. maculatum, E. angulosum var. maculatum, E. angulosum var. rectipes
Name authority A. Heller: Muhlenbergia 2: 188. (1906) Porter ex S. Watson: Proc. Amer. Acad. Arts 12: 263. (1877)
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