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angle stermed buckwheat, spotted buckwheat, spotted wild buckwheat

Havard's buckwheat, Havard's wild buckwheat

Habit Herbs, erect to spreading, annual, 1–2(–3) dm, tomentose, greenish to reddish. Herbs, erect, not scapose, (2–)3–6 × 1.5–4 dm, glabrous, grayish.
Stems

caudex absent;

aerial flowering stems erect, not striated or angled, solid, not fistulose, 0.1–0.5 dm, tomentose.

spreading, without persistent leaf bases, up to 1/5 height of plant;

caudex stems absent;

aerial flowering stems erect, slender, solid, not fistulose, 0.5–2.5 dm, glabrous, tomentose among leaves.

Leaves

basal and cauline;

basal: petiole 0.3–1 cm, floccose, blade lanceolate to obovate, 1–3(–4) × 1–1.5(–2) cm, tomentose abaxially, sparsely floccose to glabrate and grayish to greenish adaxially, margins entire or infrequently crenulate;

cauline sessile, blade lanceolate to oblanceolate, 0.5–2 × 0.3–1 cm, similar to basal blade.

basal, 1 per node;

petiole 0.5–1.5(–2.5) cm, tomentose;

blade oblanceolate to elliptic, 1–3(–5) × 0.2–1(–1.3) cm, densely white- or silvery-tomentose on both surfaces, margins plane.

Inflorescences

cymose, open, 5–25 × 10–30 cm;

branches tomentose;

bracts 3, scalelike, 0.5–2.5 × 1–2 mm.

cymose, open, divided 3–10 times, 10–40 × 10–30 cm;

branches dichotomous, glabrous;

bracts 3, scalelike, triangular, 1–2 mm.

Peduncles

spreading, straight or nearly so, filiform, (0.5–)1–3 cm, glandular-puberulent.

erect, slender, 0.5–6 cm.

Involucres

campanulate, 1–1.5(–2) × 1.5–3(–3.5) mm, glandular-puberulent;

teeth 5, erect, 0.4–0.8 mm.

1 per node, campanulate, 1.5–2.5 × (1.5–)2–3 mm, tomentose;

teeth 5, erect, 0.5–0.8 mm.

Flowers

1–2.5 mm;

perianth white to yellow, becoming pink or red, with a conspicuous rose-purple spot on each outer tepal, glandular-puberulent;

tepals dimorphic, those of outer whorl elliptic to roundish or obovate, inflated from base to middle, those of inner whorl lanceolate;

stamens included, 1–1.5(–2) mm;

filaments pilose proximally.

2.5–3 mm;

perianth yellow, densely white-pubescent;

tepals connate proximal 1/3, monomorphic, lanceolate;

stamens exserted, 2.5–3.5 mm;

filaments glabrous.

Achenes

brown, 3-gonous, 1–1.5 mm, glabrous.

brown, 2–2.5 mm, glabrous.

2n

= 40.

= 40.

Eriogonum maculatum

Eriogonum havardii

Phenology Flowering Apr–Nov. Flowering May–Sep.
Habitat Sandy to gravelly or clayey flats and slopes, mixed grassland, saltbush, creosote bush, and sagebrush communities, pinyon-juniper and montane conifer woodlands Gravelly (often calcareous) or shaley to clayey or gypsum flats and outcrops, mixed grassland, creosote, and mesquite communities, juniper woodlands
Elevation 100-2500 m (300-8200 ft) (400-)800-1800(-2100) m ((1300-)2600-5900(-6900) ft)
Distribution
from FNA
AZ; CA; ID; NM; NV; OR; UT; WA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
NM; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Eriogonum maculatum is the most common and widespread expression of the E. angulosum complex, being found in Arizona, California, southwestern Idaho, Nevada, northwestern New Mexico, eastern Oregon, western Utah, and southeastern Washington. It is often common to abundant and may even be weedy, especially along roadsides intermixed with other annual wild buckwheats. The swollen bases of the outer tepals readily distinguish the species. The greatest concentrations of the spotted wild buckwheat are found in the Mojave Desert and the Great Basin. A specimen supposedly found on the “Laramie Plains” of Wyoming (Parry s.n., 1882, ISC) is discounted as to location.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eriogonum havardii is infrequent in southeastern New Mexico (Chaves, Eddy, Lea, Lincoln, Otero, and Socorro counties) and western Texas (Brewster, Culberson, El Paso, Hudspeth, Pecos, Presidio, Terrell, Val Verde, and Winkler counties). The leaf blades are demarcated by depressed lines along their entire length, appearing as a series of folds. The bright yellow of the flowers is seen only by looking at the adaxial surface of the tepals, the abaxial surface being wholly obscured by dense white hairs. The species is worthy of cultivation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 411. FNA vol. 5, p. 288.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Synonyms E. angulosum subsp. maculatum, E. angulosum var. maculatum, E. angulosum var. rectipes E. leucophyllum
Name authority A. Heller: Muhlenbergia 2: 188. (1906) S. Watson: Proc. Amer. Acad. Arts 18: 194. (1883)
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