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angle stermed buckwheat, spotted buckwheat, spotted wild buckwheat

Brandegee's buckwheat, Brandegee's wild buckwheat

Habit Herbs, erect to spreading, annual, 1–2(–3) dm, tomentose, greenish to reddish. Herbs, spreading, sometimes scapose, 1–2.5 × (0.5–)1–2 dm, tomentose to floccose, grayish.
Stems

caudex absent;

aerial flowering stems erect, not striated or angled, solid, not fistulose, 0.1–0.5 dm, tomentose.

spreading, with persistent leaf bases, up to 1/4 height of plant;

caudex stems matted, floccose or glabrous;

aerial flowering stems erect, slender, solid, not fistulose, 1–2.5 dm, tomentose to floccose.

Leaves

basal and cauline;

basal: petiole 0.3–1 cm, floccose, blade lanceolate to obovate, 1–3(–4) × 1–1.5(–2) cm, tomentose abaxially, sparsely floccose to glabrate and grayish to greenish adaxially, margins entire or infrequently crenulate;

cauline sessile, blade lanceolate to oblanceolate, 0.5–2 × 0.3–1 cm, similar to basal blade.

strictly basal, 1 per node;

petiole 1–3(–3.5) cm, lanate;

blade oblanceolate to elliptic, 1.5–4(–5) × 0.4–1.2(–1.6) cm, densely white-tomentose adaxially, less so and greenish adaxially, margins plane.

Inflorescences

cymose, open, 5–25 × 10–30 cm;

branches tomentose;

bracts 3, scalelike, 0.5–2.5 × 1–2 mm.

capitate or umbellate-cymose, 0.1–1.5 cm wide, tomentose to floccose;

branches dichotomous, sometimes absent, tomentose to floccose;

bracts 3, scalelike, triangular, and 2–5 mm, or leaflike, lanceolate, and 1–3 × 0.2–0.7 cm.

Peduncles

spreading, straight or nearly so, filiform, (0.5–)1–3 cm, glandular-puberulent.

absent.

Involucres

campanulate, 1–1.5(–2) × 1.5–3(–3.5) mm, glandular-puberulent;

teeth 5, erect, 0.4–0.8 mm.

4–8 per cluster, turbinate, 3.5–5 × 3–4 mm, floccose or glabrous;

teeth 5, erect, 0.4–0.8 mm.

Flowers

1–2.5 mm;

perianth white to yellow, becoming pink or red, with a conspicuous rose-purple spot on each outer tepal, glandular-puberulent;

tepals dimorphic, those of outer whorl elliptic to roundish or obovate, inflated from base to middle, those of inner whorl lanceolate;

stamens included, 1–1.5(–2) mm;

filaments pilose proximally.

(2.5–)3–3.5 mm;

perianth ochroleucous, glabrous;

tepals connate proximal 1/4, monomorphic, oblanceolate to oblong;

stamens exserted, 3–3.5 mm;

filaments pilose basally.

Achenes

brown, 3-gonous, 1–1.5 mm, glabrous.

light brown, 3–3.5 mm, glabrous.

2n

= 40.

Eriogonum maculatum

Eriogonum brandegeei

Phenology Flowering Apr–Nov. Flowering Jul–Oct.
Habitat Sandy to gravelly or clayey flats and slopes, mixed grassland, saltbush, creosote bush, and sagebrush communities, pinyon-juniper and montane conifer woodlands Clay slopes and washes, sagebrush communities, juniper woodlands
Elevation 100-2500 m (300-8200 ft) 1800-2600 m (5900-8500 ft)
Distribution
from FNA
AZ; CA; ID; NM; NV; OR; UT; WA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
CO
[BONAP county map]
Discussion

Eriogonum maculatum is the most common and widespread expression of the E. angulosum complex, being found in Arizona, California, southwestern Idaho, Nevada, northwestern New Mexico, eastern Oregon, western Utah, and southeastern Washington. It is often common to abundant and may even be weedy, especially along roadsides intermixed with other annual wild buckwheats. The swollen bases of the outer tepals readily distinguish the species. The greatest concentrations of the spotted wild buckwheat are found in the Mojave Desert and the Great Basin. A specimen supposedly found on the “Laramie Plains” of Wyoming (Parry s.n., 1882, ISC) is discounted as to location.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Eriogonum brandegeei is a rare and localized species known from nine occurrences along the Arkansas River in Chaffee and Fremont counties. It is regarded as a “sensitive” species in Colorado, by both the state and the U.S. Forest Service. The Droney Gulch Natural Area has been established, in part, to protect the species. Brandegee’s wild buckwheat is related to E. brevicaule var. laxifolium, but well isolated from that taxon. An undated E. L. Greene specimen (ISC) supposedly was gathered in northern New Mexico, but that is most unlikely. A G. W. Letterman collection (MO) purportedly from Colorado Springs certainly is mislabeled.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 411. FNA vol. 5, p. 269.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Synonyms E. angulosum subsp. maculatum, E. angulosum var. maculatum, E. angulosum var. rectipes E. spathulatum var. brandegeei
Name authority A. Heller: Muhlenbergia 2: 188. (1906) Rydberg: Fl. Rocky Mts., 220, 1061. (1917)
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