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fine branch wild buckwheat, sand buckwheat, sand wild buckwheat
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Blue Mountain buckwheat, strict buckwheat
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| Shrubs, erect to spreading, not scapose, (2–)3–10(–12) × 5–15(–20) dm, white-tomentose to floccose, or green and nearly glabrous. |
Herbs, erect, loosely to densely matted, not scapose, 1–5 × 1–10 dm, tomentose or glabrous. |
spreading, often without persistent leaf bases, up to 1/2 height of plant; caudex stems absent or spreading in moving sand; aerial flowering stems spreading, slender, solid, not fistulose, 0.3–1 dm, white-tomentose, floccose to glabrate or glabrous. |
spreading to erect, with or without persistent leaf bases, up to 1/4 height of plant; caudex stems absent or spreading to matted; aerial flowering stems erect to spreading, slender, solid, not fistulose, 1–3 dm, tomentose or glabrous. |
cauline, 1 per node; petiole 0.2–0.5 cm, tomentose to floccose; blade linear-lanceolate to linear-oblanceolate to narrowly oblong, or lanceolate to narrowly elliptic, 1.5–4 × 0.2–0.8(–1.2) cm, densely white-tomentose abaxially, less so and greenish adaxially, margins plane, infrequently revolute. |
basal, 1 per node; petiole rarely twisted or curled, 1–6 cm, mostly tomentose; blade elliptic to ovate, 0.5–2.5(–4) × (0.3–)0.5–1.5 cm, lanate, tomentose to floccose on both surfaces, sometimes sparsely tomentose to floccose and greenish or floccose to subglabrous or glabrous adaxially, margins plane. |
cymose, open, 5–40 × 10–50 cm; branches dichtomous proximally, often with involucres racemosely arranged proximally, tomentose to floccose or glabrate, occasionally glabrous; bracts 3, scalelike, linear to triangular, 1–3(–6) mm. |
umbellate-cymose to cymose, 1–20 × 3–25 cm; branches dichotomous, tomentose to floccose or less often glabrous; bracts 3, scalelike, triangular, 1–3 mm. |
absent. |
absent. |
1 per node, turbinate to turbinate-campanulate, 1.5–3 × 1–2 mm, tomentose to floccose or glabrous; teeth 5, erect, 0.4–0.7 mm. |
1 per node, rarely 2–5 per cluster, narrowly turbinate to turbinate-campanulate, 4–6 × 1.5–5 mm, tomentose or glabrous; teeth 5, erect, 0.5–1.3 mm. |
(2–)2.5–3.5 mm; perianth white or pale yellow to yellow, glabrous; tepals connate proximal 1/4–1/3, essentially monomorphic, oblong to broadly obovate; stamens slightly exserted, 2–4 mm; filaments sparsely pilose proximally. |
3–5(–6) mm; perianth yellow or white to rose or purple, glabrous; tepals connate proximally, dimorphic, those of outer whorl elliptic to nearly orbiculate, 2–3 × 2–3 mm, those of inner whorl oblanceolate to oblong, 3–4 × 1–2 mm; stamens included to slightly exserted, 2–5 mm; filaments pilose proximally. |
light brown, 2.5–3.5 mm, glabrous except for minutely papillate beak. |
light brown to brown, 3–3.5 mm, glabrous. |
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AZ; CO; NM; UT
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CA; ID; MT; NV; OR; WA
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Varieties 3 (3 in the flora). Eriogonum leptocladon is localized on moving sands on the Colorado Plateau. It is not unusual, therefore, for portions of the shrub to be buried in sand, especially the main, woody trunk from which the flowering stems arise. In the Four Corners area three species share portions of overlapping ranges, and herbarium material from there can be difficult to separate. In northwestern New Mexico, var. ramosissimum approaches E. effusum, and in eastern San Juan County the two can be distinguished only by the open, spreading inflorescence branching pattern of var. ramosissimum as compared to the densely branched, compact inflorescence of E. effusum. In the field, the former is confined to sandy areas whereas the latter is typically on heavier, usually clayey soils. In south-central Utah, var. ramosissimum approaches E. nummulare. There, both are found on moving sands, and only the degree of stoutness of the branches can be used to differentiate them when cauline leaves are absent. For the most part, the inflorescence branches of E. nummulare are more rigid and stouter than those of var. ramosissimum, which tend to be flexible and slender. When leaves are present, those of the former are broader and more apically rounded compared to those of the latter. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 4 (4 in the flora). Eriogonum strictum, E. niveum, and E. ovalifolium form a complex of closely related species differing in leaf, inflorescence branching, and flower features. Variety proliferum appears to be the basal entity of the complex, approaching both E. niveum and E. ovalifolium var. pansum in its pubescence and branching pattern. Also, specimens of var. proliferum are sometimes difficult to differentiate from E. nudum var. oblongifolium. Careful observation, though, will permit well-made collections to be easily distributed among the individual species. An alternative taxonomy is to reduce all of the taxa to E. ovalifolium and recognize a series of subspecies and varieties. It is possible that additional study will show that E. strictum is sufficiently distinct from its tomentose to floccose counterparts to justify recognition of E. proliferum. In that case, both var. anserinum and var. greenei would be assigned to the latter species. Or, one could follow C. L. Hitchcock et al. (1955–1969, vol. 2) and recognize subsp. strictum as distinct from subsp. proliferum, with the latter consisting of varieties proliferum, anserinum, and greenei. Members of the Eriogonum strictum are food plants for the Bauer’s dotted-blue butterfly (Euphilotes baueri). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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1. Inflorescence branches glabrous | var. strictum |
1. Inflorescence branches tomentose to floccose | → 2 |
| var. anserinum |
2. Perianths white to rose or purple | → 3 |
3. Leaf blades grayish-tomentose to floccose on both surfaces, or greenish-tomentose to floccose adaxially | var. proliferum |
3. Leaf blades densely whitish-lanate to tomentose on both surfaces | var. greenei |
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| FNA vol. 5, p. 249. |
FNA vol. 5, p. 317. |
| Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
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| E. effusum subsp. leptocladon |
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| Torrey & A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 2(1): 129. (1857) |
Bentham: Trans. Linn. Soc. London 17: 414. (1836) |
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