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Antelope sage, James' buckwheat, James' wild buckwheat

ladder buckwheat, ladder wild buckwheat

Habit Herbs or subshrubs, compact or spreading, matted, 0.5–2.5 × 3–15 dm, tomentose to floccose. Herbs, erect, annual, (0.5–)3–6(–10) dm, glabrous, glaucous, grayish.
Stems

caudex absent or spreading;

aerial flowering stems erect, slender, solid, not fistulose, usually arising directly from a taproot, 0.5–1.5 dm, tomentose to floccose.

caudex absent;

aerial flowering stems erect, solid, not fistulose, (0.5–)1–2 dm, glabrous.

Leaves

basal, typically not in rosettes;

petiole 0.5–6 cm, tomentose to floccose;

blade usually narrowly elliptic, 1–3(–3.5) × (0.3–)0.5–1(–1.2) cm, densely tomentose abaxially, thinly tomentose, floccose or glabrous and grayish to greenish adaxially, margins entire, plane or undulate and crisped.

basal;

petiole 1–10 cm, tomentose;

blade sub-cordate to orbiculate, (1.5–)2–5(–8) × (1.5–)2–5(–8) cm, densely white-tomentose abaxially, floccose to subglabrous and green adaxially, margins often wavy.

Inflorescences

umbellate or compound-umbellate, 10–30 × 10–25 cm;

branches tomentose to floccose;

bracts 3–9, semileaflike at proximal node, 0.5–2 × 0.2–1 cm, often scalelike distally.

cymose, spreading when immature, quickly becoming narrowly erect and strict with whiplike branches, (5–)10–50(–80) × 10–50 cm;

branches glabrous, glaucous;

bracts 3, scalelike, 1–1.5(–2) × 0.4–1 mm.

Peduncles

absent or erect, straight, slender, 0.1–0.2 cm, glabrous.

Involucres

1 per node, turbinate, 1.5–7 × 2–5 mm, tomentose to floccose;

teeth 5–8, erect, 0.1–0.5 mm.

turbinate, 2–2.5(–3) × 1.5–2.5 mm, glabrous;

teeth 5, erect, 0.5–1.5 mm.

Flowers

3–8 mm, including 0.7–2 mm stipelike base;

perianth white to cream, densely pubescent abaxially;

tepals dimorphic, those of outer whorl lanceolate to elliptic, 2–5 × 1–3 mm, those of inner whorl lanceolate to fan-shaped, 1.5–6 × 2–4 mm;

stamens exserted, 2–4 mm;

filaments pilose proximally.

1.5–2 mm;

perianth white with green or reddish midribs, becoming pinkish, glabrous;

tepals dimorphic, those of outer whorl oblong, those of inner whorl lanceolate;

stamens exserted, 1.5–2.5 mm;

filaments pilose proximally.

Achenes

light brown to brown, 4–5 mm, glabrous except for sparsely pubescent beak.

dark brown to blackish, 3-gonous, 2–2.5 mm, glabrous.

2n

= 40.

Eriogonum jamesii

Eriogonum exaltatum

Phenology Flowering May–Oct.
Habitat Sandy to gravelly flats and slopes, saltbush, creosote bush, greasewood, blackbrush, and mesquite communities, rarely in pinyon-juniper woodlands
Elevation 500-1400 m (1600-4600 ft)
Distribution
from FNA
AZ; CO; KS; NM; OK; TX; including Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NV; UT
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Eriogonum jamesii is a nectar source for the rare Spalding dotted-blue butterfly (Euphilotes spaldingi).

Eriogonum jamesii and E. arcuatum (see below) are considered “life medicines” and used ceremonially by Native Americans (C. Arnold, pers. comm.; A. B. Reagan 1929; P. A. Vestal 1952).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eriogonum insigne has been a troublesome taxon.  The type, collected by E. Palmer in 1876, is a curious specimen from near Paragonah in Iron County, Utah.  That expression has not been recollected and, after another frustrating review of the type material, it is now referred to E. deflexum, along with other odd specimens from southern Nevada and California that have always been difficult to assign.  As a result, the tall, upright plants with long, whiplike branches of northwestern Arizona (Mohave County), southern Nevada (northeastern Clark and southern Lincoln counties), and southwestern Utah (southwestern Washington County), are now recognized under the name E. exaltatum.  Some specimens remain problematic.  Immature plants of E. exaltatum (Goodding 2302, GH, MIN, MO, NY, RM, UC) resemble E. bifurcatum. Specimens from Eureka Valley (DeDecker 4741, NY) and western Pahrump Valley (Abrams 14248, DS, GH, NY) in Inyo County, California, appear to belong to E. exaltatum.

Like Eriogonum exaltatum, some populations of E. deflexum var. deflexum have nearly erect involucres.  The plants from the Buried Hills (Nye and Clark counties, Nevada) are particularly odd.  Single plants from a few collections in Kane County, Utah, have sessile, seemingly erect involucres in the forks of inflorescence branches.  None of these plants has the characteristic long, whiplike branches of E. exaltatum, and they are not included in E. deflexum var. deflexum. The type of E. deflexum var. rectum from San Bernardino County, California (Reveal & Broome 6385, CAS, NY, US, etc.), has individual specimens with both deflexed and somewhat erect involucres, along with whiplike branches.  Collections from Imperial and San Diego counties, previously associated with what is here termed E. exaltatum, are now considered specimens of E. deflexum.  Clearly, more work is required to understand these species fully.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf margins undulate, frequently crisped; flowers 3-5(-6) mm; se Arizona, s New Mexico, sw Texas
var. undulatum
1. Leaf margins plane, not crisped; flowers 4-8 mm; e Arizona, c and s Colorado, wc Kansas, New Mexico, w Oklahoma, n and w Texas
→ 2
2. Inflorescences compound-umbellate; e Arizona, c and s Colorado, New Mexico, w Oklahoma, n and w Texas
var. jamesii
2. Inflorescences simple-umbellate; wc Kansas
var. simplex
Source FNA vol. 5, p. 367. FNA vol. 5, p. 397.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Oligogonum Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. jamesii var. jamesii, E. jamesii var. simplex, E. jamesii var. undulatum
Name authority Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 14: 7. (1856) M. E. Jones: Contr. W. Bot. 15: 61. (1929)
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