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Antelope sage, James' buckwheat, James' wild buckwheat

spreading buckwheat, spreading wild buckwheat

Habit Herbs or subshrubs, compact or spreading, matted, 0.5–2.5 × 3–15 dm, tomentose to floccose. Shrubs, spreading, not scapose, (1.5–)2–5(–7) × 5–15 dm, grayish- to reddish-brown-tomentose to floccose and gray or, rarely, thinly floccose and greenish.
Stems

caudex absent or spreading;

aerial flowering stems erect, slender, solid, not fistulose, usually arising directly from a taproot, 0.5–1.5 dm, tomentose to floccose.

spreading to erect, typically without persistent leaf bases, up to 1/2 height of plant;

caudex stems absent;

aerial flowering stems erect to spreading, slender, solid, not fistulose, 0.3–0.8 dm, floccose or glabrous.

Leaves

basal, typically not in rosettes;

petiole 0.5–6 cm, tomentose to floccose;

blade usually narrowly elliptic, 1–3(–3.5) × (0.3–)0.5–1(–1.2) cm, densely tomentose abaxially, thinly tomentose, floccose or glabrous and grayish to greenish adaxially, margins entire, plane or undulate and crisped.

cauline, 1 per node;

petiole 0.2–0.7 cm, tomentose to floccose;

blade oblanceolate to oblong or obovate, (1–)1.5–3 × (0.2–)0.3–0.7 cm, densely white-tomentose abaxially, white-floccose to glabrate or green and glabrous adaxially, margins plane.

Inflorescences

umbellate or compound-umbellate, 10–30 × 10–25 cm;

branches tomentose to floccose;

bracts 3–9, semileaflike at proximal node, 0.5–2 × 0.2–1 cm, often scalelike distally.

cymose, 10–30(–40) × 10–40 cm;

branches dichotomous, white-floccose to glabrate or subglabrous;

bracts 3, scalelike, triangular, 0.5–2(–5) mm.

Peduncles

absent or mostly erect, slender, 0.3–2.5 cm, floccose.

Involucres

1 per node, turbinate, 1.5–7 × 2–5 mm, tomentose to floccose;

teeth 5–8, erect, 0.1–0.5 mm.

1 per node, turbinate, 1.5–2.5(–3) × 1–2 mm, tomentose to floccose;

teeth 5, erect, 0.3–0.6 mm.

Flowers

3–8 mm, including 0.7–2 mm stipelike base;

perianth white to cream, densely pubescent abaxially;

tepals dimorphic, those of outer whorl lanceolate to elliptic, 2–5 × 1–3 mm, those of inner whorl lanceolate to fan-shaped, 1.5–6 × 2–4 mm;

stamens exserted, 2–4 mm;

filaments pilose proximally.

2–4 mm;

perianth yellow, glabrous;

tepals connate proximal 1/4, essentially monomorphic, elliptic to obovate;

stamens mostly exserted, 2–4.5 mm;

filaments sparsely pilose proximally.

Achenes

light brown to brown, 4–5 mm, glabrous except for sparsely pubescent beak.

brown, 2–2.5 mm, glabrous.

Eriogonum jamesii

Eriogonum effusum

Phenology Flowering Jun–Sep.
Habitat Sandy to rocky slopes and flats, mixed grassland and sagebrush communities, juniper and montane conifer woodlands
Elevation 1200-2500 m (3900-8200 ft)
Distribution
from FNA
AZ; CO; KS; NM; OK; TX; including Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; MT; NE; NM; SD; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Eriogonum jamesii is a nectar source for the rare Spalding dotted-blue butterfly (Euphilotes spaldingi).

Eriogonum jamesii and E. arcuatum (see below) are considered “life medicines” and used ceremonially by Native Americans (C. Arnold, pers. comm.; A. B. Reagan 1929; P. A. Vestal 1952).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eriogonum effusum is rather common on the northern Great Plains and along the eastern slope of the Rocky Mountains in central and eastern Colorado, southeastern Montana, western Nebraska, northern New Mexico, southwestern South Dakota, and southeastern Wyoming. Some specimens from Chaffee County, Colorado, are thinly floccose and greenish (Atwood & Welsh 29689, BRY) and thus similar to E. leptocladon. A collection from Pyramid Lake, Washoe County, Nevada (Frandsen & Brown 182, NESH) is clearly mislabeled. A roadside collection of E. effusum gathered near Little America, Sweetwater County, Wyoming, in 1961 (G. Mason 4025, ASU) was an introduction that has not persisted.

The spreading wild buckwheat is occasionally merged with Eriogonum microthecum even though the two are morphologically distinct and their ranges do not overlap. Plants in New Mexico are sometimes difficult to distinguish from the related E. leptocladon var. ramosissimum. The species is the food plant for the Rita dotted-blue butterfly (Euphilotes rita). A hybrid between Eriogonum effusum and E. pauciflorum has been named E. ×nebraskense Rydberg [E. multiceps Nees subsp. nebraskense (Rydberg) S. Stokes; E. pauciflorum Pursh var. nebraskense (Rydberg) Reveal]. The hybrid is known from Weld County, Colorado; Cheyenne and Kimball counties, Nebraska; and Converse and Platte counties, Wyoming.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf margins undulate, frequently crisped; flowers 3-5(-6) mm; se Arizona, s New Mexico, sw Texas
var. undulatum
1. Leaf margins plane, not crisped; flowers 4-8 mm; e Arizona, c and s Colorado, wc Kansas, New Mexico, w Oklahoma, n and w Texas
→ 2
2. Inflorescences compound-umbellate; e Arizona, c and s Colorado, New Mexico, w Oklahoma, n and w Texas
var. jamesii
2. Inflorescences simple-umbellate; wc Kansas
var. simplex
Source FNA vol. 5, p. 367. FNA vol. 5, p. 248.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Oligogonum Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. jamesii var. jamesii, E. jamesii var. simplex, E. jamesii var. undulatum
Synonyms E. microthecum var. effusum
Name authority Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 14: 7. (1856) Nuttall: Proc. Acad. Nat. Sci. Philadelphia 4: 15. (1848)
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