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bottle stopper, desert trumpet, Indian pipeweed

fraziers well buckwheat, Ripley's wild buckwheat

Habit Herbs, erect, perennial, occasionally flowering first year, 1–10(–15) dm, glabrous, usually glaucous, grayish. Subshrubs, sprawling to decumbent, occasionally scapose, 0.5–2.5(–4) × 0.5–5(–7) dm, grayish-tomentose.
Stems

caudex compact;

aerial flowering stems erect, solid or hollow and fistulose, (0.2–)2–5 dm, glabrous, usually glaucous, occasionally hirsute proximally.

decumbent to slightly spreading or erect, often with persistent leaf bases, up to 1/4 height of plant;

caudex stems absent or compact;

aerial flowering stems spreading to erect, slender, solid, not fistulose, 0.05–2.5(–3.5) dm, tomentose.

Leaves

basal;

petiole 2–6 cm, hirsute;

blade oblong-ovate to oblong or rounded to reniform, (0.5–)1–2.5(–3) × (0.5–)1–2(–2.5) cm, short-hirsute and grayish or greenish on both surfaces, sometimes less so or glabrous and green adaxially, margins occasionally undulate.

cauline, 1 per node or fasciculate;

petiole 0.05–0.15 cm, villous;

blade narrowly oblanceolate, 0.2–0.6 × 0.05–0.1 cm, densely white-tomentose abaxially, thinly floccose to villous adaxially, margins revolute.

Inflorescences

cymose, open, spreading to erect, 5–70 × 5–50 cm;

branches occasionally fistulose, glabrous, usually glaucous;

bracts 3, scalelike, 1–2.5(–5) × 1–2.5 mm.

cymose-umbellate or capitate and reduced to single involucre;

branches usually dichotomous, sometimes with secondaries suppressed, occasionally absent, tomentose;

bracts absent.

Peduncles

erect, straight, filiform to capillary, 0.5–2(–3.5) cm, glabrous.

erect, 0.1–1.5 cm, thinly floccose.

Involucres

turbinate, 1–1.5 × 1–1.8 mm, glabrous;

teeth 5, erect, 0.4–0.6 mm.

1 per node, campanulate, 3–3.5 × 3–3.5 mm, thinly floccose or villous to subglabrous;

teeth 3–5, erect, 0.7–1 mm.

Flowers

(1–)2–3(–4) mm;

perianth yellow with greenish or reddish midribs, densely hirsute with coarse curved hairs;

tepals monomorphic, narrowly ovoid to ovate;

stamens exserted, 1.3–2.5 mm;

filaments glabrous or sparsely pubescent proximally.

3.5–4.5 mm;

perianth white, glabrous;

tepals connate proximal 1/5, dimorphic, those of outer whorl nearly orbiculate, 3–3.5 mm wide, those of inner whorl broadly obovate, 2–2.5 mm wide;

stamens long-exserted, 4–5 mm;

filaments villous to densely pilose proximally.

Achenes

light brown to brown, lenticular to 3-gonous, 2–2.5 mm, glabrous.

light brown, 2–2.5 mm, glabrous.

2n

= 32.

Eriogonum inflatum

Eriogonum ripleyi

Phenology Flowering year-round. Flowering Mar–Jun.
Habitat Sandy to gravelly washes, flats, and slopes, mixed grassland, saltbush, creosote bush, mesquite, and sagebrush communities, pinyon and/or juniper woodlands Sandy clay flats and slopes on edge of sandstone outcrops, oak-juniper woodlands
Elevation -30-1800(-2000) m (-100-5900(-6600) ft) 1000-1900 m (3300-6200 ft)
Distribution
from FNA
AZ; CA; CO; NM; NV; UT; Mexico (Baja California, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ
[BONAP county map]
Discussion

The cause of the fistulose stem and inflorescence branches in Eriogonum inflatum was imaginatively attributed by A. M. Stone and C. T. Mason (1979) to the larvae of gall insects. This fallacy continues to appear in the literature. Greenhouse studies have shown that stems of this and some other species of the genus inflate without the presence of any insects. Other researchers have shown that the inflation involves a build-up of CO2 within the stems, which take over as the primary photosynthetic body as leaves wilt or eventually dry up and fall away from the plant (C. D. Osmond et al. 1987). Not all individuals of E. inflatum will have fistulose stems and branches, as this feature is partly a function of available moisture: the drier the conditions, the less pronounced the inflation. Stems produced in the summer tend to be inflated less frequently than those produced in the spring.

The “annual” phase of Eriogonum inflatum is distinct from its truly annual relatives. Its flowering stems and inflorescence branches are distinctly grayish, whereas those of the true annuals are green or yellowish green.

As circumscribed here, Eriogonum inflatum occurs in Arizona, southern and east-central California, western Colorado, northwestern New Mexico, central and southern Nevada, and southern and eastern Utah.

Some Native Americans occasionally ate newly emerged stems of Eriogonum inflatum (S. A. Weber and P. D. Seaman 1985; M. L. Zigmond 1981). The hollow stems were used as drinking tubes (Weber and Seaman) and pipes (E. W. Gifford 1936). This wild buckwheat is a food plant for the desert metalmark butterfly (Apodemia mormo deserti).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Eriogonum ripleyi is known only from two areas in Arizona, one near Frazier’s Well in Coconino County and a second in the Verdi Valley area of southeastern Yavapai and extreme northwestern Maricopa counties. The species is worthy of cultivation. Ripley’s wild buckwheat is considered a “sensitive” species in Arizona.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 386. FNA vol. 5, p. 253.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Synonyms E. glaucum, E. inflatum var. deflatum, E. trichopes subsp. glaucum
Name authority Torrey & Frémont: in J. C. Frémont, Rep. Exped. Rocky Mts., 317. (1845) J. T. Howell: Leafl. W. Bot. 4: 5. (1944)
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