Heerman buckwheat, Heermann's buckwheat, Heermann's wild buckwheat
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bastard-sage, bastard-sage buckwheat, Wright's buckwheat, Wright's wild buckwheat
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Shrubs and subshrubs, spreading to rounded and occasionally erect, not scapose, (0.5–)1–20 × 2–25 dm, glabrous or occasionally floccose, sometimes scabrellous, greenish, infrequently grayish. |
Shrubs, subshrubs, or herbs, rarely scapose, (1–)1.5–10 × 1–15(–18) dm or, if matted, 0.1–2.5(–3) × 0.5–3(–5) dm, , lanate to thinly tomentose, or glabrous, grayish to greenish or reddish. |
spreading or erect, without persistent leaf bases, up to 1/2 height of plant; caudex stems absent; aerial flowering stems erect or nearly so, slender to stout, solid, not fistulose, 0.02–0.5 dm, thinly tomentose or glabrous. |
spreading to erect, with or without persistent leaf bases, up to 1/2 or more height of plant; caudex stems absent or spreading, occasionally matted; aerial flowering stems erect to spreading, stout to slender, solid, not fistulose, (0.1–)0.5–4(–6) dm, tomentose, floccose, or glabrous. |
cauline, 1 per node, quickly deciduous; petiole 0.1–1.5 cm, floccose or glabrous; blade linear, oblanceolate or spatulate or elliptic, or oblong, (0.4–)1–2(–4) × 0.1–0.8 cm, tomentose to floccose or glabrous abaxially, floccose to thinly floccose or glabrous adaxially, margins plane. |
basal and fasciculate in terminal tufts, or cauline and fasciculate, occasionally 1 per node; petiole 0.02–0.5(–1) cm, tomentose to floccose; blade oblanceolate to broadly elliptic, 0.1–3 × 0.1–1 cm, tomentose to floccose, sometimes subglabrous or glabrous and green adaxially, margins plane, sometimes revolute. |
cymose or racemose, 1–25(–30) × 1–30(–35) cm; branches dichotomous, sometimes with secondaries suppressed, smooth or angled to ridged and grooved, glabrous or occasionally floccose or scabrous; bracts 3, scalelike, 0.3–2 mm. |
virgate or cymose with involucres disposed at tips racemosely arranged involucres, rarely capitate, (1–)5–20 × (1–)10–40 cm; branches dichotomous, tomentose, floccose, or glabrous; bracts 3, triangular, scalelike, 0.5–3.5 mm. |
absent. |
absent. |
1 per node, narrowly turbinate or campanulate, 0.7–3 × 0.7–4 mm, glabrous, infrequently floccose; teeth 5, erect, 0.3–0.7 mm. |
1 per node, turbinate to narrowly campanulate, (0.7–)1–4 × 1–2.5 mm, tomentose, floccose, or glabrous; teeth 5, erect, 0.3–1 mm. |
(1.5–)2–4 mm; perianth white, yellowish white, pink, or reddish, glabrous; tepals connate proximal 1/4, dimorphic, those of outer whorl obovate to orbiculate, those of inner whorl narrowly lanceolate to oblong; stamens exserted, 2–5 mm; filaments pilose proximally. |
1–4 mm; perianth white to pink or rose, glabrous; tepals connate proximal 1/4, monomorphic, obovate; stamens exserted, 1.5–4 mm; filaments glabrous or sparsely pilose proximally. |
light brown to brown, 2–5 mm, glabrous. |
light brown to brown, (1–)1.5–3 mm, glabrous. |
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AZ; CA; NV; UT
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AZ; CA; NM; NV; TX; UT; including nw Mexico
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Varieties 8 (8 in the flora). As Eriogonum heermannii is here circumscribed, the number of varieties is dramatically decreased from past presentations, with E. apachense reduced to synonymy under a now greatly expanded var. argense. Also included in that variety is the more stoutly branched var. subracemosum. The southern var. heermannii of basically desert ranges and the more northern var. occidentale of the Coast Ranges in California are maintained, but their separation is more traditional than certain. The fragile and bulky nature of many dried, often poorly prepared specimens, and the tendency for leaves to fall away have made varietal identification within E. heermannii difficult. Eriogonum heermannii varieties are food plants for Ellis’s dotted-blue butterfly (Euphilotes ellisi) and the Mormon metalmark (Apodemia mormo mormo). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 9 (6 in the flora). Eriogonum wrightii is subdivided into several varieties most of which are distinct, although a few have rather indistinct boundaries. Some of the variation has yet to be fully resolved, especially in the var. nodosum complex where, at least in Mexico, one additional expression remains to be named. A clear distinction between var. subscaposum and E. kennedyi is not possible in southern California (see discussion below). Nearly all of the varieties of E. wrightii are in cultivation, although the most elegant (var. olanchense) has yet to be so honored. Few ethnobotanical uses of bastard-sage are reported in the literature. L. C. Wyman and S. K. Harris (1951) noted that the Kayenta Navajo use it (var. wrightii) as an emetic, while M. L. Zigmond (1981) stated that the Kawaiisu used the pounded seeds (probably of var. subscaposum) in a beverage or as a dry meal. Members of the species are food plants for the rare Rita dotted-blue butterfly (Euphilotes rita), the Pacific dotted-blue (E. enoptes), the veined blue (Plebeius neurona), and the Mormon metalmark (Apodemia mormo mormo). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Flowering stems and inflorescence branches round or angled, scabrellous or papillate-scabrous; plants often densely branched | → 2 |
1. Flowering stems and inflorescence branches round, not angled, usually smooth, glabrous or thinly tomentose; plants sparsely branched | → 3 |
2. Flowering stems and inflorescence branches round, usually distinctly scabrellous or infrequently papillate-scabrous; Arizona, se California, Nevada | var. argense |
2. Flowering stems and inflorescence branches sharply ridged and deeply grooved, minutely scabrellous; nw Arizona, se California, s Nevada, sw Utah | var. sulcatum |
3. Involucres not racemosely arranged on inflorescence branches, or only last 2-3 so disposed; inflorescences diffusely branched, glabrous; Great Basin or northeast edge of Mojave Desert | → 4 |
3. Involucres racemosely arranged along inflorescence branches or at least at tips of branches; inflorescences openly branched, glabrous or floccose to thinly tomentose; Mojave and Sonoran deserts | → 5 |
4. Inflorescence branches mostly slender, smooth, not ridged, not spinose; inflorescences 3-15(-23) × 5-20 cm; subshrubs or shrubs, 3-7 × 5-12(-15) dm; Great Basin, ec California and Nevada | var. humilius |
4. Inflorescence branches stoutish, faintly ridged and grooved, spinose; inflorescences 3-7(-10) × 3-10(-12) cm; shrubs, 1-3 × 1.5-5(-8) dm; Mojave Desert, sw Utah and nw Arizona | var. subspinosum |
5. Inflorescence branches thinly tomentose to floccose; s California, s Nevada, nw Arizona | var. floccosum |
5. Inflorescence branches glabrous; s Nevada, California | → 6 |
6. Involucres racemosely arranged; inflorescence branches whiplike; s Nevada | var. clokeyi |
6. Involucres racemosely arranged only distally; inflorescence branches not whiplike; California | → 7 |
7. Leaf blades 0.5-1.5 cm, glabrous abaxially; inflorescence branches stout; sc California | var. heermannii |
7. Leaf blades 1.5-3(-4) cm, mostly tomentose to floccose abaxially (at least in early anthesis); inflorescence branches slender; sw California | var. occidentale |
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1. Plants loosely to compactly matted or herbs | → 2 |
1. Plants shrubs or subshrubs | → 3 |
2. Plants 0.5-2.5(-3) dm; leaf blades 0.5-1(-1.2) cm; involucres 1.5-4 mm; e and s California,wc Nevada | var. subscaposum |
2. Plants 0.1-0.3(-0.6) dm; leaf blades 0.1-0.25 cm; involucres 0.8-1.7(-2) mm; se California | var. olanchense |
3. Flowering stems and branches grayish, lanate to densely tomentose; sw Arizona, se California | var. nodosum |
3. Flowering stems and inflorescence branches whitish, reddish, or greenish, tomentose to floccose; widespread | → 4 |
4. Petiole bases forming distinct ring around stem; leaf blades 0.2-0.6(-1) × 0.1-0.3(-0.4) cm; s California | var. membranaceum |
4. Petiole bases not forming distinct ring around stem; leaf blades 0.5-3 × 0.2-1 cm; sw United States, Mexico | → 5 |
5. Leaf blades 0.5-1.5 × 0.2-0.5(-0.7) cm; involucres 2-2.5 mm; flowers 2.5-3.5 mm; se California to w Texas | var. wrightii |
5. Leaf blades 1.5-3 × 0.5-1 cm; involucres 3-4 mm; flowers 3-4 mm; c and nw California | var. trachygonum |
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FNA vol. 5, p. 304. |
FNA vol. 5, p. 289. |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. zionis |
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E. geniculatum |
E. trachygonum subsp. wrightii |
Durand & Hilgard: in War Department [U.S.], Pacif. Railr. Rep. 5(3): 14. (1857) |
Torrey ex Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 14: 15. (1856) |
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