Heerman buckwheat, Heermann's buckwheat, Heermann's wild buckwheat
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cushion buckwheat, cushion wild buckwheat, oval-leaf buckwheat, oval-leaf eriogonum
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Shrubs and subshrubs, spreading to rounded and occasionally erect, not scapose, (0.5–)1–20 × 2–25 dm, glabrous or occasionally floccose, sometimes scabrellous, greenish, infrequently grayish. |
Herbs, forming pulvinate to cespitose maps, usually scapose, 0.2–3 × 0.5–5 dm, floccose to tomentose or lanate. |
spreading or erect, without persistent leaf bases, up to 1/2 height of plant; caudex stems absent; aerial flowering stems erect or nearly so, slender to stout, solid, not fistulose, 0.02–0.5 dm, thinly tomentose or glabrous. |
decumbent to spreading, with persistent leaf bases, up to 1/5 height of plant; caudex stems matted to spreading; aerial flowering stems scapelike, spreading to erect, infrequently decumbent to ascending, slender, solid, not fistulose, 0.03–4 dm, floccose to tomentose or lanate or nearly glabrous. |
cauline, 1 per node, quickly deciduous; petiole 0.1–1.5 cm, floccose or glabrous; blade linear, oblanceolate or spatulate or elliptic, or oblong, (0.4–)1–2(–4) × 0.1–0.8 cm, tomentose to floccose or glabrous abaxially, floccose to thinly floccose or glabrous adaxially, margins plane. |
basal, 1 per node; petiole not twisted or curled, 0.1–10 cm, mostly tomentose; blade oblanceolate to elliptic or spatulate to rounded, 0.2–6 × (0.1–)0.2–1.5 cm, lanate to tomentose or floccose, sometimes less so adaxially, margins plane, occasionally brownish. |
cymose or racemose, 1–25(–30) × 1–30(–35) cm; branches dichotomous, sometimes with secondaries suppressed, smooth or angled to ridged and grooved, glabrous or occasionally floccose or scabrous; bracts 3, scalelike, 0.3–2 mm. |
capitate or rarely umbellate, 0.7–5(–7) × 1.5–5 cm; bracts 3, scalelike, linear to triangular, 0.8–4 mm. |
absent. |
absent. |
1 per node, narrowly turbinate or campanulate, 0.7–3 × 0.7–4 mm, glabrous, infrequently floccose; teeth 5, erect, 0.3–0.7 mm. |
1 per node or (2–)3–15 per cluster, turbinate to turbinate-campanulate, (2–)3.5–5(–8) × 2–4 mm, tomentose to floccose; teeth 5, erect, 0.1–1 mm. |
(1.5–)2–4 mm; perianth white, yellowish white, pink, or reddish, glabrous; tepals connate proximal 1/4, dimorphic, those of outer whorl obovate to orbiculate, those of inner whorl narrowly lanceolate to oblong; stamens exserted, 2–5 mm; filaments pilose proximally. |
(2.5–)3–6(–7) mm; perianth yellow or white to cream, rose, red, or purple, glabrous; tepals connate proximally, dimorphic, those of outer whorl usually oval to orbiculate, 2–4 × 2–4 mm, those of inner whorl oblanceolate to elliptic, 3–7 × 0.8–1.5 mm; stamens mostly included, 1–3 mm; filaments pilose proximally. |
light brown to brown, 2–5 mm, glabrous. |
light brown to brown, 2–3 mm, glabrous. |
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AZ; CA; NV; UT
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AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC
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Varieties 8 (8 in the flora). As Eriogonum heermannii is here circumscribed, the number of varieties is dramatically decreased from past presentations, with E. apachense reduced to synonymy under a now greatly expanded var. argense. Also included in that variety is the more stoutly branched var. subracemosum. The southern var. heermannii of basically desert ranges and the more northern var. occidentale of the Coast Ranges in California are maintained, but their separation is more traditional than certain. The fragile and bulky nature of many dried, often poorly prepared specimens, and the tendency for leaves to fall away have made varietal identification within E. heermannii difficult. Eriogonum heermannii varieties are food plants for Ellis’s dotted-blue butterfly (Euphilotes ellisi) and the Mormon metalmark (Apodemia mormo mormo). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 11 (11 in the flora). Eriogonum ovalifolium is a highly diverse and widespread complex of generally distinct but sometimes intergrading varieties. Several varieties are in cultivation and make worthy additions, especially to the rock garden. The dimorphic nature of the tepals is obvious only in fully mature flowers. In some populations of the more depauperate varieties, such as var. nivale and var. depressum, the tepals may not be as distinctly dimorphic. Nonetheless, the overall aspect of the species is unmistakable. There are several reports of traditional use of Eriogonum ovalifolium by Native Americans. P. Train et al. (1941) indicated that a decoction of the roots is used in Nevada for colds. R. V. Chamberlin (1911) reported that the Gosiute Indians in southwestern Utah used it in a poultice or wash to treat venereal diseases. Members of the species are food plants for Bauer’s dotted-blue butterfly (Euphilotes baueri). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Flowering stems and inflorescence branches round or angled, scabrellous or papillate-scabrous; plants often densely branched | → 2 |
1. Flowering stems and inflorescence branches round, not angled, usually smooth, glabrous or thinly tomentose; plants sparsely branched | → 3 |
2. Flowering stems and inflorescence branches round, usually distinctly scabrellous or infrequently papillate-scabrous; Arizona, se California, Nevada | var. argense |
2. Flowering stems and inflorescence branches sharply ridged and deeply grooved, minutely scabrellous; nw Arizona, se California, s Nevada, sw Utah | var. sulcatum |
3. Involucres not racemosely arranged on inflorescence branches, or only last 2-3 so disposed; inflorescences diffusely branched, glabrous; Great Basin or northeast edge of Mojave Desert | → 4 |
3. Involucres racemosely arranged along inflorescence branches or at least at tips of branches; inflorescences openly branched, glabrous or floccose to thinly tomentose; Mojave and Sonoran deserts | → 5 |
4. Inflorescence branches mostly slender, smooth, not ridged, not spinose; inflorescences 3-15(-23) × 5-20 cm; subshrubs or shrubs, 3-7 × 5-12(-15) dm; Great Basin, ec California and Nevada | var. humilius |
4. Inflorescence branches stoutish, faintly ridged and grooved, spinose; inflorescences 3-7(-10) × 3-10(-12) cm; shrubs, 1-3 × 1.5-5(-8) dm; Mojave Desert, sw Utah and nw Arizona | var. subspinosum |
5. Inflorescence branches thinly tomentose to floccose; s California, s Nevada, nw Arizona | var. floccosum |
5. Inflorescence branches glabrous; s Nevada, California | → 6 |
6. Involucres racemosely arranged; inflorescence branches whiplike; s Nevada | var. clokeyi |
6. Involucres racemosely arranged only distally; inflorescence branches not whiplike; California | → 7 |
7. Leaf blades 0.5-1.5 cm, glabrous abaxially; inflorescence branches stout; sc California | var. heermannii |
7. Leaf blades 1.5-3(-4) cm, mostly tomentose to floccose abaxially (at least in early anthesis); inflorescence branches slender; sw California | var. occidentale |
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1. Leaf blades 0.2-1.2(-2) cm; scapes usually 0.3-5(-9) cm, rarely longer; involucres usually 2-4.5 (rarely 5-8) mm | → 2 |
1. Leaf blades usually 1-6 cm, occasionally shorter; scapes (1-)5-30(-40) cm; involucres (3.5-)4-6.5(-8) mm | → 8 |
2. Perianths yellow; c Nevada and ec California | var. caelestinum |
2. Perianths white, sometimes rose, purple, or red; widespread | → 3 |
| → 4 |
3. Leaf margins not brownish or, if so, plants of high elevation, central Sierra Nevada | → 5 |
4. Leaf blades densely lanate, margins brownish | var. eximium |
4. Leaf blades tomentose, margins not brownish | var. williamsiae |
| → 6 |
| → 7 |
6. Leaf blades white-lanate; San Bernardino Mountains, s California | var. vineum |
6. Leaf blades tomentose to floccose; s Sierra Nevada, California | var. monarchense |
7. Leaf blades greenish and thinly tomentose at least adaxially, usually elliptic, infrequently oblong to spatulate; scapes often suberect to decumbent, usually thinly floccose; Rocky Mountains of Alberta, Idaho, Montana, and Wyoming, desert ranges of ne Nevada and e Oregon | var. depressum |
7. Leaves densely lanate to white-tomentose on both surfaces or only slightly less so adaxially, not at all greenish, usually round; scapes usually erect, lanate or tomentose; Sierra-Cascade cordillera, desert ranges of Great Basin | var. nivale |
8. Scapes usually 1-5(-7.5) cm; leaf margins brownish; rare, wc Nevada, ec California | → 9 |
8. Scapes usually (4-)7-30(-40) cm, rarely shorter; leaf margins only rarely brownish; widespread | → 10 |
9. Leaf blades densely lanate, brownish margins | var. eximium |
9. Leaf blades white-tomentose, margins not brownish | var. williamsiae |
10. Inflorescences umbellate, branches 1-3 cm; c Idaho and Montana | var. pansum |
10. Inflorescences capitate, branches absent; widespread | → 11 |
11. Flowers 5-7 mm; involucres 5-7 mm; s California | var. vineum |
11. Flowers 4-5 mm; involucres 4-6.5 mm; widespread | → 12 |
| var. ovalifolium |
12. Perianths white to rose or purple. | → 13 |
13. Leaf blades spatulate, oblong, or obovate to oval; scapes (4-)5-20 cm; widespread | var. purpureum |
13. Leaf blades oblanceolate to narrowly elliptic; scapes (15-)20-30(-40) cm; Montana and Wyoming | var. ochroleucum |
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FNA vol. 5, p. 304. |
FNA vol. 5, p. 319. |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
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E. ovalifolium var. caelestinum, E. ovalifolium var. depressum, E. ovalifolium var. eximium, E. ovalifolium var. monarchense, E. ovalifolium var. nivale, E. ovalifolium var. ochroleucum, E. ovalifolium var. ovalifolium, E. ovalifolium var. pansum, E. ovalifolium var. purpureum, E. ovalifolium var. vineum, E. ovalifolium var. williamsiae |
E. geniculatum |
Eucycla ovalifolia |
Durand & Hilgard: in War Department [U.S.], Pacif. Railr. Rep. 5(3): 14. (1857) |
Nuttall: J. Acad. Nat. Sci. Philadelphia 7: 50, plate 8, fig. 1. (1834) |
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