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Heerman buckwheat, Heermann's buckwheat, Heermann's wild buckwheat

Fort Mohave wild buckwheat, Fort Mojave buckwheat

Habit Shrubs and subshrubs, spreading to rounded and occasionally erect, not scapose, (0.5–)1–20 × 2–25 dm, glabrous or occasionally floccose, sometimes scabrellous, greenish, infrequently grayish. Herbs, erect, annual, (0.5–)1–7 dm, glabrous and sparsely floccose, greenish.
Stems

spreading or erect, without persistent leaf bases, up to 1/2 height of plant;

caudex stems absent;

aerial flowering stems erect or nearly so, slender to stout, solid, not fistulose, 0.02–0.5 dm, thinly tomentose or glabrous.

caudex absent;

aerial flowering stems erect, solid, not fistulose, (0.3–) 0.7–3 dm, thinly floccose or glabrous, floccose proximally.

Leaves

cauline, 1 per node, quickly deciduous;

petiole 0.1–1.5 cm, floccose or glabrous;

blade linear, oblanceolate or spatulate or elliptic, or oblong, (0.4–)1–2(–4) × 0.1–0.8 cm, tomentose to floccose or glabrous abaxially, floccose to thinly floccose or glabrous adaxially, margins plane.

basal, occasionally cauline;

basal: petiole 2–6(–10) cm, floccose, blade oblong-oblanceolate to obovate, (1.5–)2–8 × (0.8–)1–3 cm, thinly floccose or glabrous and green on both surfaces, margins entire;

cauline: petiole 0.5–3 cm, thinly floccose, blade elliptic to obovate, 0.7–3 × 0.2–2 cm, similar to basal blade.

Inflorescences

cymose or racemose, 1–25(–30) × 1–30(–35) cm;

branches dichotomous, sometimes with secondaries suppressed, smooth or angled to ridged and grooved, glabrous or occasionally floccose or scabrous;

bracts 3, scalelike, 0.3–2 mm.

paniculate, open to diffuse, (5–)10–50 × 5–50 cm;

branches not fistulose, glabrous except for floccose nodes and proximal branches;

bracts 3, scalelike, 0.5–3 × 0.3–1 mm.

Peduncles

absent.

erect, straight, capillary, 0.5–2 cm, glabrous or thinly floccose.

Involucres

1 per node, narrowly turbinate or campanulate, 0.7–3 × 0.7–4 mm, glabrous, infrequently floccose;

teeth 5, erect, 0.3–0.7 mm.

narrowly turbinate to turbinate, 1–1.5(–1.8) × 0.6–1.2 mm, glabrous;

teeth 4, erect, 0.2–0.5 mm.

Flowers

(1.5–)2–4 mm;

perianth white, yellowish white, pink, or reddish, glabrous;

tepals connate proximal 1/4, dimorphic, those of outer whorl obovate to orbiculate, those of inner whorl narrowly lanceolate to oblong;

stamens exserted, 2–5 mm;

filaments pilose proximally.

1–2.5(–3) mm;

perianth white with greenish or reddish midribs to pale yellow with greenish midribs, becoming pink to reddish, densely short-villous;

tepals monomorphic, oblong to narrowly ovate;

stamens exserted, 1–1.5 mm;

filaments glabrous.

Achenes

light brown to brown, 2–5 mm, glabrous.

dark brown to black, 3-gonous, 1.8–2 mm, glabrous.

Eriogonum heermannii

Eriogonum ordii

Phenology Flowering Mar–Jul.
Habitat Gravelly to clayey flats and slopes, mixed grassland communities, oak and conifer woodlands
Elevation 200-1400 m (700-4600 ft)
Distribution
from FNA
AZ; CA; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 8 (8 in the flora).

As Eriogonum heermannii is here circumscribed, the number of varieties is dramatically decreased from past presentations, with E. apachense reduced to synonymy under a now greatly expanded var. argense. Also included in that variety is the more stoutly branched var. subracemosum. The southern var. heermannii of basically desert ranges and the more northern var. occidentale of the Coast Ranges in California are maintained, but their separation is more traditional than certain. The fragile and bulky nature of many dried, often poorly prepared specimens, and the tendency for leaves to fall away have made varietal identification within E. heermannii difficult.

Eriogonum heermannii varieties are food plants for Ellis’s dotted-blue butterfly (Euphilotes ellisi) and the Mormon metalmark (Apodemia mormo mormo).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eriogonum ordii is infrequently encountered (rarely locally common) along the inner Coast Ranges from Monterey and San Benito counties south through Fresno, Merced, and San Luis Obispo counties to Ventura County, then eastward across northern Los Angeles County to the hills east of Bakersfield in Kern County. This distribution is based on confirmed modern collections. The type (J. G. Lemmon 4189, ASU, BM, DS, G, GH, ISC, K, P, UC, US) supposedly was collected near Fort Mohave, Mohave County, Arizona, in 1884. Two T. Brandegee specimens reportedly were gathered on the boundary of San Diego and Imperial counties, one at Split Mountain (Apr 1905, UC) and the second along San Felipe Creek (4 Apr 1901, UC). Brandegee’s label data often are dubious, and these disjunct sites are discounted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Flowering stems and inflorescence branches round or angled, scabrellous or papillate-scabrous; plants often densely branched
→ 2
1. Flowering stems and inflorescence branches round, not angled, usually smooth, glabrous or thinly tomentose; plants sparsely branched
→ 3
2. Flowering stems and inflorescence branches round, usually distinctly scabrellous or infrequently papillate-scabrous; Arizona, se California, Nevada
var. argense
2. Flowering stems and inflorescence branches sharply ridged and deeply grooved, minutely scabrellous; nw Arizona, se California, s Nevada, sw Utah
var. sulcatum
3. Involucres not racemosely arranged on inflorescence branches, or only last 2-3 so disposed; inflorescences diffusely branched, glabrous; Great Basin or northeast edge of Mojave Desert
→ 4
3. Involucres racemosely arranged along inflorescence branches or at least at tips of branches; inflorescences openly branched, glabrous or floccose to thinly tomentose; Mojave and Sonoran deserts
→ 5
4. Inflorescence branches mostly slender, smooth, not ridged, not spinose; inflorescences 3-15(-23) × 5-20 cm; subshrubs or shrubs, 3-7 × 5-12(-15) dm; Great Basin, ec California and Nevada
var. humilius
4. Inflorescence branches stoutish, faintly ridged and grooved, spinose; inflorescences 3-7(-10) × 3-10(-12) cm; shrubs, 1-3 × 1.5-5(-8) dm; Mojave Desert, sw Utah and nw Arizona
var. subspinosum
5. Inflorescence branches thinly tomentose to floccose; s California, s Nevada, nw Arizona
var. floccosum
5. Inflorescence branches glabrous; s Nevada, California
→ 6
6. Involucres racemosely arranged; inflorescence branches whiplike; s Nevada
var. clokeyi
6. Involucres racemosely arranged only distally; inflorescence branches not whiplike; California
→ 7
7. Leaf blades 0.5-1.5 cm, glabrous abaxially; inflorescence branches stout; sc California
var. heermannii
7. Leaf blades 1.5-3(-4) cm, mostly tomentose to floccose abaxially (at least in early anthesis); inflorescence branches slender; sw California
var. occidentale
Source FNA vol. 5, p. 304. FNA vol. 5, p. 390.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. heermannii var. argense, E. heermannii var. clokeyi, E. heermannii var. floccosum, E. heermannii var. heermannii, E. heermannii var. humilius, E. heermannii var. occidentale, E. heermannii var. subspinosum, E. heermannii var. sulcatum
Synonyms E. geniculatum E. tenuissimum
Name authority Durand & Hilgard: in War Department [U.S.], Pacif. Railr. Rep. 5(3): 14. (1857) S. Watson: Proc. Amer. Acad. Arts 21: 468. (1886)
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