California buckwheat, California wild buckwheat, eastern Mojave buckwheat, eastern Mojave wild buckwheat
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bare-stem buckwheat, barestem wild buckwheat, naked buckwheat, naked wild buckwheat, nude buckwheat
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Shrubs or subshrubs, compact to spreading or rounded and more or less erect, occasionally decumbent, infrequently scapose, (1–)2–15 × 2–25(–30) dm, tomentose to canescent, floccose, or glabrous. |
Herbs, mostly erect, infrequently scapose, (0.5–)1–15(–20) × 0.5–3 dm, glabrous or floccose to tomentose, usually greenish, occasionally grayish. |
sprawling or spreading to erect, often with persistent leaf bases, up to 1/2 or more height of plant; caudex stems absent or matted to spreading; aerial flowering stems erect to spreading, slender, solid, not fistulose, 0.3–2.5(–3) dm, tomentose, canescent, or glabrous. |
spreading to erect, with or without persistent leaf bases, up to 1/4 height of plant; caudex stems absent; aerial flowering stems erect to spreading, slender to stout, solid or hollow, occasionally fistulose, 0.3–4(–10) dm, glabrous or lanate to tomentose or floccose. |
cauline, 1 per node or fasciculate; petiole 0.1–0.3 cm, canescent; blade linear to linear-oblanceolate or oblanceolate, 0.6–1.5(–1.8) × 0.05–0.4(–0.6) cm, white-tomentose or canescent to subglabrous abaxially, tomentose or canescent and grayish, subglabrous, or glabrous and green adaxially, margins often revolute. |
basal or sheathing up stem 0.5–4 dm; petiole 1–10 cm, glabrous or tomentose; blade oblanceolate to elliptic or ovate, 1–6 × (0.3–)1–4 cm, densely white-lanate or tomentose abaxially, tomentose to floccose or subglabrous to glabrous adaxially, margins plane or undulate-crisped. |
cymose, infrequently cymose-umbellate or capitate, compact to open, occasionally flat-topped, 0.2–20 × 0.2–15 cm; branches dichotomous, infrequently absent, tomentose to canescent or glabrous; bracts usually 3, scalelike, triangular, and 1–3 mm, or leaflike, linear to oblanceolate, and 3–10 × 1–3 mm. |
cymose, rarely umbellate or capitate, 2–100(–150) × 2–40(–80) cm; branches usually dichotomous, glabrous or tomentose to floccose or sparsely pubescent; bracts 3, scalelike, 0.5–3(–5) mm. |
absent. |
absent. |
(1–)3–8 per cluster, turbinate to campanulate, 2–4 × 1.5–3 mm, canescent, pubescent, glabrous, or subglabrous; teeth 5, erect, 0.3–1.2 mm. |
1 per node or 2–10 per cluster, turbinate to turbinate-campanulate, (2.5–)3–5(–7) × (1.5–)2–4 mm, glabrous, tomentose, or sparsely pubescent; teeth 5–8, 0.2–0.6 mm. |
2.5–3 mm; perianth white to pinkish, glabrous or pubescent; tepals connate proximal 1/4, monomorphic, usually elliptic to obovate; stamens exserted, 2.5–5 mm; filaments subglabrous or pubescent proximally. |
(1.5–)2–4 mm; perianth white or yellow, sometimes pink or rose, glabrous or pubescent; tepals connate proximal 1/4, monomorphic, oblong to obovate; stamens exserted, 2–5 mm; filaments pilose proximally. |
light brown to brown, 1.8–2.5 mm, glabrous. |
light brown to brown, 1.5–3.5 mm, glabrous. |
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AZ; CA; NV; UT; including nw Mexico
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CA; NV; OR; WA; nw Mexico
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Varieties 5 (4 in the flora). Eriogonum fasciculatum is a complex, polyploid series of variants that are generally distinct but often difficult to distinguish morphologically. Variety emphereium Reveal is confined to central Baja California, Mexico. The introduction of Eriogonum fasciculatum as a decorative roadside plant by the California Department of Transportation is resulting in hybrid populations involving E. cinereum. The aggressively weedy and (for Arizona) exotic variety foliosum is rapidly invading the native habitat of var. polifolium. Members of E. fasciculatum are food plants for several butterflies, notably the Bernardino dotted-blue (Euphilotes bernardino), lupine blue (Plebeius lupini), Mormon metalmark (Apodemia mormo), and Behr’s metalmark (A. virgulti). Probably the butterfly most commonly seen with the species is the nut-brown hairstreak (Satyrium saepium), which frequents plants in full flower. Eriogonum fasciculatum is also the most important native source of honey in California. This widespread species was used extensively by Native Americans for a variety of ailments. Its application for pain and headaches (D. P. Barrows 1900; K. Hedges 1986; E. W. Voegelin 1938) was rather common, as was its general use for diarrhea (Hedges; Voegelin). M. L. Zigmond (1981) reported that the Kawaiisu lined their acorn granaries with leaves of var. proliferum to keep out rain—a daunting challenge given the size of the leaves! L. Hinton (1975) reported the use of a decoction of dried flowers and roots to maintain a healthy heart, and M. C. Stevenson (1915) indicated that a powder derived from the roots was used by the Zuñi to treat wounds, whereas a root decoction was taken for colds and hoarseness. B. R. Bocek (1984) reported that the Costanoan Indians of California used a decoction of the plant to treat unspecified urinary problems. F. H. Elmore (1943) reported the use of a decoction of var. proliferum by the Navajo (Diné) people as an anti-witchcraft medicine. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 13 (13 in the flora). Several local groups of Native Americans in California used members of this species in a variety of ways. S. A. Barrett and E. W. Gifford (1933) and S. M. Schenck and E. W. Gifford (1952) reported the consumption of raw young stems that are rather moist and tasty, although there is a sour aftertaste. The Kawaiisu used the hollow stems (probably var. westonii, rather than var. pauciflorum) as drinking tubes and as pipes (M. L. Zigmond 1981). Zigmond reported also that the roots of var. pauciflorum are used as an infusion for coughs. Members of Eriogonum nudum are food plants for the Bauer’s dotted-blue butterfly (Euphilotes baueri), the Pacific dotted-blue (E. enoptes), the gorgon copper (Gaeides gorgon), and the Mormon metalmark (Apodemia mormo). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Leaves thinly white-tomentose abaxially, glabrous adaxially; perianths and involucres mostly glabrous; plants usually decumbent; primarily coastal 80c. Eriogonum fasciculatum var. fasciculatum | → 1 |
1. Leaves, perianths, and involucres pubescent or, if subglabrous, then plants of desert regions; plants erect to rounded; generally inland | → 2 |
2. Leaves light yellow-green, subglabrous adaxially; involucres and perianths glabrous or subglabrous | var. flavoviride |
2. Leaves dark green or grayish, usually pubescent adaxially; involucres and perianths pubescent | → 3 |
3. Inflorescences capitate to cymose-umbellate, rarely cymose; leaves canescent on both surfaces or densely tomentose abaxially and canescent adaxially; leaf margins plane or infrequently revolute; mainly deserts | var. polifolium |
3. Inflorescences open and mostly cymose; leaves densely white-tomentose abaxially, less so to green and floccose adaxially; leaf margins usually tightly revolute; mainly inner coastal foothills | var. foliolosum |
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1. Involucres and inflorescence branches tomentose to floccose; leaves usually basal | → 2 |
1. Involucres and inflorescence branches glabrous or, if pubescent, leaves sheathing up stems | → 3 |
2. Perianths white; flowering stems lanate to tomentose; leaf blades 2-3.5 cm; s Sierra Nevada | var. regirivum |
2. Perianths yellow or white to rose; flowering stems tomentose to floccose; leaf blades 2-4 cm; s Oregon, n California, wc Nevada | var. oblongifolium |
3. Leaves sheathing up stems, margins often strongly undulate-crisped | → 4 |
3. Leaves basal, margins plane or slightly undulate-crisped | → 8 |
4. Flowering stems tomentose | → 5 |
4. Flowering stems glabrous | → 6 |
5. Perianths pubescent; flowers 1.5-2 mm; involucres 3-4 mm; Sierra Nevada, California | var. regirivum |
5. Perianths glabrous; flowers 3-4 mm; involucres 4-6 mm; Coast Ranges, California | var. decurrens |
6. Leaf blades densely lanate abaxially, tomentose adaxially; involucres 5-10 per cluster; Sierra Nevada, California | var. murinum |
6. Leaf blades tomentose abaxially, less so to floccose, glabrous, or nearly so adaxially; involucres 1 per node or 2-5 per cluster; widespread, Coast Ranges, California | → 7 |
7. Flowering stems occasionally fistulose; involucres (2-)3-5 per cluster; perianths white to pink, rarely yellowish | var. auriculatum |
7. Flowering stems strongly fistulose; involucres 1 per node; perianths pale yellowish white to yellow or white | var. indictum |
8. Involucres 1(-2) per cluster | var. westonii |
8. Involucres 2-10 per cluster | → 12 |
9. Flowering stems fistulose; perianths yellow, infrequently white; c California | var. westonii |
9. Flowering stems not fistulose or, if so, plants of sw California; perianths white, rarely yellow | → 10 |
10. Perianths pubescent or, infrequently, glabrous; s California | var. pauciflorum |
10. Flowers glabrous; sw Oregon, n California, Nevada | → 11 |
11. Leaf blades 1-5 cm; sw Oregon, n California (including w slope of Sierra Nevada), Washington | var. nudum |
11. Leaf blades 1-2 cm; Sierra Nevada of California and adjacent desert ranges of wc Nevada | var. deductum |
12. Perianths pubescent, often yellow | → 13 |
12. Perianths usually glabrous abaxially, white, rarely yellow | → 14 |
13. Flowering stems not fistulose; leaf blades floccose or glabrous adaxially, margins plane; plants of nonaridregions | var. pubiflorum |
13. Flowering stems slightly to distinctly fistulose; leaf blades tomentose to floccose adaxially, margins undulate-crisped; plants of arid regions | var. westonii |
14. Inflorescences capitate or nearly so; alpine, Sierra Nevada | E. nudumvar. scapigerum |
14. Inflorescences cymose or, if capitate, not alpine. | → 15 |
15. Involucres 1(-2) per cluster, 5- 7 mm; mountains of s California | var. pauciflorum |
15. Involucres 2-10 per cluster, 3-5 mm; mountains and foothills of c and n California or coastal bluffs. [16. Shifted to left margin—Ed.] | → 16 |
16. Inflorescences cymose and branched 2 or more times; involucres 2-5 per cluster; mountains and foothills of c and n California, Oregon, and Washington. | var. nudum |
16. Inflorescences capitate or cymose and branched 1-2 times; involucres 5-10 per cluster; coastal bluffs, sw Oregon, ne California | var. paralinum |
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FNA vol. 5, p. 297. |
FNA vol. 5, p. 309. |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
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E. nudum var. auriculatum, E. nudum var. decurrens, E. nudum var. deductum, E. nudum var. indictum, E. nudum var. murinum, E. nudum var. nudum, E. nudum var. oblongifolium, E. nudum var. paralinum, E. nudum var. pauciflorum, E. nudum var. pubiflorum, E. nudum var. regirivum, E. nudum var. westonii, E. nudumvar. scapigerum |
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E. latifolium subsp. nudum |
Bentham: Trans. Linn. Soc. London 17: 411. (1836) |
Douglas ex Bentham: Trans. Linn. Soc. London 17: 413. (1836) |
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