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California buckwheat, California wild buckwheat, eastern Mojave buckwheat, eastern Mojave wild buckwheat

slender buckwheat, slender wild buckwheat

Habit Shrubs or subshrubs, compact to spreading or rounded and more or less erect, occasionally decumbent, infrequently scapose, (1–)2–15 × 2–25(–30) dm, tomentose to canescent, floccose, or glabrous. Subshrubs or shrubs, erect to spreading, not scapose, 0.2–1.5 × (0.6–)1–13(–16) dm, white- to tannish-tomentose, floccose, or glabrous.
Stems

sprawling or spreading to erect, often with persistent leaf bases, up to 1/2 or more height of plant;

caudex stems absent or matted to spreading;

aerial flowering stems erect to spreading, slender, solid, not fistulose, 0.3–2.5(–3) dm, tomentose, canescent, or glabrous.

spreading to erect, typically without persistent leaf bases, up to 1/2 height of plant;

caudex stems absent or spreading;

aerial flowering stems erect to spreading, slender, solid, not fistulose, 0.05–1.5 dm, lanate, tomentose, floccose, subglabrous, or glabrous.

Leaves

cauline, 1 per node or fasciculate;

petiole 0.1–0.3 cm, canescent;

blade linear to linear-oblanceolate or oblanceolate, 0.6–1.5(–1.8) × 0.05–0.4(–0.6) cm, white-tomentose or canescent to subglabrous abaxially, tomentose or canescent and grayish, subglabrous, or glabrous and green adaxially, margins often revolute.

cauline, 1 per node or fasciculate;

petiole 0.1–0.5 cm, tomentose to floccose or glabrous;

blade usually elliptic, sometimes linear to obovate, 0.3–3.5 × (0.07–)0.1–1.2 cm, tomentose abaxially, less so or glabrous adaxially, margins occasionally revolute.

Inflorescences

cymose, infrequently cymose-umbellate or capitate, compact to open, occasionally flat-topped, 0.2–20 × 0.2–15 cm;

branches dichotomous, infrequently absent, tomentose to canescent or glabrous;

bracts usually 3, scalelike, triangular, and 1–3 mm, or leaflike, linear to oblanceolate, and 3–10 × 1–3 mm.

cymose, compact, often flat-topped, 0.5–6(–12) × 1–10(–13) cm;

branches dichotomous, whitish-lanate to brownish- or reddish-tomentose to floccose or glabrate, infrequently green or gray and subglabrous or glabrous;

bracts 3, scalelike, linear to triangular, 1–5 mm.

Peduncles

absent.

absent or mostly erect, slender, 0.3–1.5 cm, tomentose to floccose.

Involucres

(1–)3–8 per cluster, turbinate to campanulate, 2–4 × 1.5–3 mm, canescent, pubescent, glabrous, or subglabrous;

teeth 5, erect, 0.3–1.2 mm.

1 per node, turbinate, (1.5–)2–3.5(–4) × 1.3–2.5(–3) mm, tomentose, floccose, subglabrous, or glabrous;

teeth 5, erect, (0.3–)0.5–1(–1.7) mm.

Flowers

2.5–3 mm;

perianth white to pinkish, glabrous or pubescent;

tepals connate proximal 1/4, monomorphic, usually elliptic to obovate;

stamens exserted, 2.5–5 mm;

filaments subglabrous or pubescent proximally.

1.5–3(–4) mm;

perianth yellow or white to pink, orange, rose, red, or occasionally cream, glabrous;

tepals connate proximal 1/5–2/5, essentially monomorphic, oblong to obovate;

stamens usually exserted, 2.5–4 mm;

filaments sparsely to densely puberulent proximally.

Achenes

light brown to brown, 1.8–2.5 mm, glabrous.

brown, 1.5–3 mm, glabrous.

Eriogonum fasciculatum

Eriogonum microthecum

Distribution
from FNA
AZ; CA; NV; UT; including nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 5 (4 in the flora).

Eriogonum fasciculatum is a complex, polyploid series of variants that are generally distinct but often difficult to distinguish morphologically. Variety emphereium Reveal is confined to central Baja California, Mexico. The introduction of Eriogonum fasciculatum as a decorative roadside plant by the California Department of Transportation is resulting in hybrid populations involving E. cinereum. The aggressively weedy and (for Arizona) exotic variety foliosum is rapidly invading the native habitat of var. polifolium. Members of E. fasciculatum are food plants for several butterflies, notably the Bernardino dotted-blue (Euphilotes bernardino), lupine blue (Plebeius lupini), Mormon metalmark (Apodemia mormo), and Behr’s metalmark (A. virgulti). Probably the butterfly most commonly seen with the species is the nut-brown hairstreak (Satyrium saepium), which frequents plants in full flower. Eriogonum fasciculatum is also the most important native source of honey in California.

This widespread species was used extensively by Native Americans for a variety of ailments. Its application for pain and headaches (D. P. Barrows 1900; K. Hedges 1986; E. W. Voegelin 1938) was rather common, as was its general use for diarrhea (Hedges; Voegelin). M. L. Zigmond (1981) reported that the Kawaiisu lined their acorn granaries with leaves of var. proliferum to keep out rain—a daunting challenge given the size of the leaves! L. Hinton (1975) reported the use of a decoction of dried flowers and roots to maintain a healthy heart, and M. C. Stevenson (1915) indicated that a powder derived from the roots was used by the Zuñi to treat wounds, whereas a root decoction was taken for colds and hoarseness. B. R. Bocek (1984) reported that the Costanoan Indians of California used a decoction of the plant to treat unspecified urinary problems. F. H. Elmore (1943) reported the use of a decoction of var. proliferum by the Navajo (Diné) people as an anti-witchcraft medicine.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 13 (13 in the flora).

Eriogonum microthecum is used as browse by deer and to a lesser degree by cattle and sheep. Some forms are now in cultivation. The species is reportedly used by the Piute of Nevada in the treatment of tuberculosis, lameness, rheumatism, and bladder trouble (P. Train et al. 1941). S. A. Weber and P. D. Seaman (1985) stated that A. F. Whiting found the plants being used as a tea by the Havasupai in northern Arizona. Members of E. microthecum are food plants for subspecies of the rare pallid blue butterfly (Euphilotes pallescens). Also found on this species is the cythera metalmark (Apodemia mormo cythera). Some authors have referred E. effusum to this species, even though the ranges of the two species do not overlap and intermediates are unknown.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves thinly white-tomentose abaxially, glabrous adaxially; perianths and involucres mostly glabrous; plants usually decumbent; primarily coastal 80c. Eriogonum fasciculatum var. fasciculatum
→ 1
1. Leaves, perianths, and involucres pubescent or, if subglabrous, then plants of desert regions; plants erect to rounded; generally inland
→ 2
2. Leaves light yellow-green, subglabrous adaxially; involucres and perianths glabrous or subglabrous
var. flavoviride
2. Leaves dark green or grayish, usually pubescent adaxially; involucres and perianths pubescent
→ 3
3. Inflorescences capitate to cymose-umbellate, rarely cymose; leaves canescent on both surfaces or densely tomentose abaxially and canescent adaxially; leaf margins plane or infrequently revolute; mainly deserts
var. polifolium
3. Inflorescences open and mostly cymose; leaves densely white-tomentose abaxially, less so to green and floccose adaxially; leaf margins usually tightly revolute; mainly inner coastal foothills
var. foliolosum
1. Perianths yellow
→ 2
1. Perianths various shades of white, cream, orange, pink, or red, not yellow
→ 4
2. Flowering stems and inflorescence branches usually glabrous; e Oregon and wc Idaho
var. microthecum
2. Flowering stems and inflorescence branches tomentose to floccose; se Oregon and sw Idaho s to e California and w Nevada
→ 3
3. Leaf blades (0.2-)0.3-0.6(-0.8) cm wide; flowers (1.5-)2-2.5(-3) mm; involucres 2-2.5 mm; achenes 1.5-2 mm; se Oregon and sw Idaho s to e California and w Nevada
var. ambiguum
3. Leaf blades 0.5-1.2 cm wide; flowers 2.5-3 mm; involucres 2.5-4 mm; achenes 2.5-3 mm; ne California, nw Nevada
var. schoolcraftii
4. Tomentum whitish (see also var. alpinum of the Sierra Nevada, California); flowering stems and inflorescence branches infrequently glabrous
→ 5
4. Tomentum brownish or reddish (may be white in var. alpinum), or flowering stems and inflorescence branches essentially glabrous
→ 6
5. Leaf margins not revolute; flowering inflorescence branches floccose or glabrous; northern phase of species
var. laxiflorum
5. Leaf margins revolute or nearly so; flowering inflorescence branches lanate to tomentose, or if subglabrous or glabrous, then southern phase of species
var. simpsonii
6. Plants shrubs, 3-6 dm
→ 7
6. Plants subshrubs, 0.2-1.5(-2) dm
→ 8
7. Flowering stems and inflorescence branches tomentose when young, becoming floccose at maturity; flowers 1.5-2(-2.5) mm; achenes 1.8-2 mm; Death Valley region, California
var. panamintense
7. Flowering stems and inflorescence branches lanate to tomentose at maturity; flowers 2-2.5(-3) mm; achenes 2.5-3 mm; Transverse Ranges, California
E. microthecumvar. corymbosoides
8. Leaf blades elliptic or ovate, margins not revolute; flowers (1.5-)2-3.5(-4) mm
→ 9
8. Leaf blades linear or linear-oblanceolate to narrowly elliptic, margins often revolute; flowers 1.5-2.5 mm
→ 10
9. Leaf blades 0.5-1 × (0.2-)0.3-0.5(-0.6) cm; involucres (2-)2.5-3 mm; flowers (2.5-)3-3.5(-4) mm; San Gabriel Mountains, California
var. johnstonii
9. Leaf blades 0.3-0.7(-0.8) × 0.1-0.4 cm; involucres 2.5-3.5 mm; flowers (1.5-)2-3.5 mm; desert ranges of se California, c Nevada, and Utah1f. Eriogonum microthecum var. lapidicola [8. Shifted to left margin—Ed.]
→ 8
10. Involucres 3-4 mm; perianths cream; San Bernardino Mountains, California
var. lacus-ursi
10. Involucres (1.5-)2-3 mm; perianths white, pink, red, or rose; Sierra Nevada of California, or e Nevada and w Utah
→ 11
11. Flowering stems glabrous; leaf blades sparsely floccose or glabrous adaxially; desert ranges, se Nevada
var. arceuthinum
11. Flowering stems white- to brownish-floccose to subglabrous, or reddish-tomentose to floccose; leaf blades floccose to subglabrous adaxially; California or wc Utah
→ 12
12. Tomentum white to brownish; Sierra Nevada, California
var. alpinum
12. Tomentum reddish; desert ranges, wc Utah
var. phoeniceum
Source FNA vol. 5, p. 297. FNA vol. 5, p. 242.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. fasciculatum var. flavoviride, E. fasciculatum var. foliolosum, E. fasciculatum var. polifolium
E. microthecum var. alpinum, E. microthecum var. ambiguum, E. microthecum var. arceuthinum, E. microthecum var. johnstonii, E. microthecum var. lacus-ursi, E. microthecum var. laxiflorum, E. microthecum var. microthecum, E. microthecum var. panamintense, E. microthecum var. phoeniceum, E. microthecum var. schoolcraftii, E. microthecum var. simpsonii, E. microthecumvar. corymbosoides
Name authority Bentham: Trans. Linn. Soc. London 17: 411. (1836) Nuttall: Proc. Acad. Nat. Sci. Philadelphia 4: 15. (1848)
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